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Polychaeta taxon details

Prionospio Malmgren, 1867

129620  (urn:lsid:marinespecies.org:taxname:129620)

accepted
Genus
Anaspio Chamberlin, 1920 (subjective synonym)
Apoprionospio Foster, 1969 (subjective synonymy)
Aquilaspio Foster, 1971 (subjective synonymy)
Kinbergella McIntosh, 1909 (subjective synonym)
Minuspio Foster, 1971 (subjective synonym)
Periptyches Grube, 1873 (subjective synonym)
Prionospio (Apoprionospio) [auctorum] (superseded combination)
Prionospio (Aquilaspio) [auctorum] (superseded combination)
Prionospio (Minuspio) [auctorum] (superseded combination)
Prionospio (Prionospio) [auctorum] (superseded combination)
Species Prionospio aluta Maciolek, 1985
Species Prionospio amarsupiata Neal & Altamira in Paterson et al., 2016
Species Prionospio andamanensis Hylleberg & Nateewathana, 1991
Species Prionospio anneae Radashevsky, 2015
Species Prionospio anuncata Fauchald, 1972
Species Prionospio aucklandica Augener, 1923
Species Prionospio austella Delgado-Blas, 2015
Species Prionospio australiensis Blake & Kudenov, 1978
Species Prionospio bocki Söderström, 1920
Species Prionospio branchilucida Altamira, Glover & Paterson in Paterson et al., 2016
Species Prionospio caribensis Delgado-Blas, 2014
Species Prionospio caspersi Laubier, 1962
Species Prionospio cerastae Radashevsky, 2015
Species Prionospio cirrifera Wirén, 1883
Species Prionospio convexa Imajima, 1990
Species Prionospio cooki Radashevsky, 2015
Species Prionospio coorilla Wilson, 1990
Species Prionospio cornuta Hylleberg & Nateewathana, 1991
Species Prionospio crassumbranchiata Delgado-Blas, 2015
Species Prionospio cristata Foster, 1971
Species Prionospio cristaventralis Delgado-Blas, Díaz-Díaz & Viéitez, 2018
Species Prionospio dayi (Foster, 1969)
Species Prionospio decipiens Söderström, 1920
Species Prionospio delta Hartman, 1965
Species Prionospio depauperata Imajima, 1990
Species Prionospio dubia Day, 1961
Species Prionospio ehlersi Fauvel, 1928
Species Prionospio elegantula Imajima, 1990
Species Prionospio elongata Imajima, 1990
Species Prionospio ergeni Dagli & Çinar, 2009
Species Prionospio fallax Söderström, 1920
Species Prionospio fauchaldi Maciolek, 1985
Species Prionospio festiva (Grube, 1872)
Species Prionospio gracilis Hartmann-Schröder, 1962
Species Prionospio grossa Imajima, 1990
Species Prionospio henriki Hylleberg & Nateewathana, 1991
Species Prionospio hermesia Neal & Paterson in Paterson et al., 2016
Species Prionospio heterobranchia Moore, 1907
Species Prionospio jamaicensis Delgado-Blas, 2014
Species Prionospio japonicus Okuda, 1935
Species Prionospio jonatani Delgado-Blas, 2015
Species Prionospio jubata Blake, 1996
Species Prionospio kaplani Altamira, Glover & Paterson in Paterson et al., 2016
Species Prionospio kirrae Wilson, 1990
Species Prionospio komaeti Hylleberg & Nateewathana, 1991
Species Prionospio krusadensis Fauvel, 1929
Species Prionospio kulin Wilson, 1990
Species Prionospio laciniosa Maciolek, 1985
Species Prionospio lighti Maciolek, 1985
Species Prionospio lineata Imajima, 1990
Species Prionospio lylei Radashevsky, 2015
Species Prionospio malayensis (Caullery, 1914)
Species Prionospio marsupiala Blake, 1996
Species Prionospio membranacea Imajima, 1990
Species Prionospio multibranchiata Berkeley, 1927
Species Prionospio multicristata Hutchings & Rainer, 1979
Species Prionospio multipinnulata (Blake & Kudenov, 1978)
Species Prionospio neenae Hylleberg & Nateewathana, 1991
Species Prionospio newportensis Reish, 1959
Species Prionospio nielseni Hylleberg & Nateewathana, 1991
Species Prionospio nirripa Wilson, 1990
Species Prionospio nonatoi Peixoto & Paiva, 2019
Species Prionospio oligopinnulata Delgado-Blas, 2015
Species Prionospio oshimensis Imajima, 1990
Species Prionospio paradisea Imajima, 1990
Species Prionospio parapari Delgado-Blas, Díaz-Díaz & Viéitez, 2018
Species Prionospio patagonica Augener, 1923
Species Prionospio paucipinnulata Blake & Kudenov, 1978
Species Prionospio perkinsi Maciolek, 1985
Species Prionospio peruana Hartmann-Schröder, 1962
Species Prionospio phuketensis Hylleberg & Nateewathana, 1991
Species Prionospio plumosa M. Sars in G.O. Sars, 1872
Species Prionospio polybranchiata Fauvel, 1929
Species Prionospio pulchra Imajima, 1990
Species Prionospio pygmaeus Hartman, 1961
Species Prionospio pyramidalis (Hutchings & Turvey, 1984)
Species Prionospio queenslandica Blake & Kudenov, 1978
Species Prionospio rosariae Delgado-Blas, 2014
Species Prionospio rotalis Mohammad, 1970
Species Prionospio rotunda Delgado-Blas, 2015
Species Prionospio rugosa Sigvaldadóttir, 1997
Species Prionospio runei Hylleberg & Nateewathana, 1991
Species Prionospio saccifera Mackie & Hartley, 1990
Species Prionospio saldanha Day, 1961
Species Prionospio sandersi Maciolek, 1981
Species Prionospio sexoculata Augener, 1918
Species Prionospio sishaensis Wu & Chen, 1964
Species Prionospio solisi Peixoto & Paiva, 2019
Species Prionospio somaliensis Cognetti-Varriale, 1988
Species Prionospio spongicola Wesenberg-Lund, 1958
Species Prionospio steenstrupi Malmgren, 1867
Species Prionospio tatura Wilson, 1990
Species Prionospio tetelensis Gibbs, 1971
Species Prionospio texana Hartman, 1951
Species Prionospio thalanji Wilson & Humphreys, 2001
Species Prionospio tridentata Blake & Kudenov, 1978
Species Prionospio tripinnata Maciolek, 1985
Species Prionospio unilamellata Sigvaldadóttir & Desbruyères, 2003
Species Prionospio vallensis Neal & Paterson in Paterson et al., 2016
Species Prionospio variegata Imajima, 1990
Species Prionospio vermillionensis Fauchald, 1972
Species Prionospio wambiri Wilson, 1990
Species Prionospio wireni Maciolek, 1985
Species Prionospio yuriel Wilson, 1990

Species Prionospio capensis McIntosh, 1885 (taxon inquirendum, indeterminable)
Species Prionospio malmgreni Claparède, 1869 (taxon inquirendum, indeterminable, probably based on a composite of several species (fide Maciolek, 1985))
Species Prionospio nova Annenkova, 1938 (taxon inquirendum, indeterminable fragment (fide Foster, 1969, 1971))

Subgenus Prionospio (Apoprionospio) [auctorum] accepted as Prionospio Malmgren, 1867 (superseded combination)
Subgenus Prionospio (Aquilaspio) [auctorum] accepted as Prionospio Malmgren, 1867 (superseded combination)
  » Species Prionospio (Aquilaspio) krusadensis Fauvel, 1929 represented as Prionospio krusadensis Fauvel, 1929 (disused subgenus rank)
  » Species Prionospio (Aquilaspio) convexa Imajima, 1990 accepted as Prionospio convexa Imajima, 1990 (superseded original combination)
  » Species Prionospio (Aquilaspio) grossa Imajima, 1990 accepted as Prionospio grossa Imajima, 1990 (superseded original combination)
  » Species Prionospio (Aquilaspio) multipinnulata Blake & Kudenov, 1978 accepted as Prionospio multipinnulata (Blake & Kudenov, 1978) (superseded original combination)
Subgenus Prionospio (Minuspio) [auctorum] accepted as Prionospio Malmgren, 1867 (superseded combination)
  » Species Prionospio (Minuspio) anatolica Dagli & Çinar, 2011
  » Species Prionospio (Minuspio) maciolekae Dagli & Çinar, 2011
  » Species Prionospio (Minuspio) sanmartini Delgado-Blas, Díaz-Díaz & Viéitez, 2019
  » Species Prionospio (Minuspio) aluta Maciolek, 1985 accepted as Prionospio aluta Maciolek, 1985 (superseded subgenus)
  » Species Prionospio (Minuspio) cirrifera Wirén, 1883 accepted as Prionospio cirrifera Wirén, 1883 (superseded subgenus)
  » Species Prionospio (Minuspio) delta Hartman, 1965 accepted as Prionospio delta Hartman, 1965 (superseded subgenus)
  » Species Prionospio (Minuspio) elegantula Imajima, 1990 accepted as Prionospio elegantula Imajima, 1990 (superseded original combination)
  » Species Prionospio (Minuspio) elongata Imajima, 1990 accepted as Prionospio elongata Imajima, 1990 (superseded combination)
  » Species Prionospio (Minuspio) fauchaldi Maciolek, 1985 accepted as Prionospio fauchaldi Maciolek, 1985 (superseded subgenus)
  » Species Prionospio (Minuspio) japonica Okuda, 1935 accepted as Prionospio japonicus Okuda, 1935 (superseded combination)
  » Species Prionospio (Minuspio) laciniosa Maciolek, 1985 accepted as Prionospio laciniosa Maciolek, 1985 (superseded subgenus)
  » Species Prionospio (Minuspio) lighti Maciolek, 1985 accepted as Prionospio lighti Maciolek, 1985 (superseded subgenus)
  » Species Prionospio (Minuspio) multibranchiata Berkeley, 1927 accepted as Prionospio multibranchiata Berkeley, 1927 (superseded subgenus)
  » Species Prionospio (Minuspio) perkinsi Maciolek, 1985 accepted as Prionospio perkinsi Maciolek, 1985 (superseded original combination)
  » Species Prionospio (Minuspio) pulchra Imajima, 1990 accepted as Prionospio pulchra Imajima, 1990 (superseded combination)
  » Species Prionospio (Minuspio) sandersi Maciolek, 1981 accepted as Prionospio sandersi Maciolek, 1981 (superseded original combination)
  » Species Prionospio (Minuspio) wireni Maciolek, 1985 accepted as Prionospio wireni Maciolek, 1985 (superseded subgenus)
Subgenus Prionospio (Paraprionospio) Caullery, 1914 accepted as Paraprionospio Caullery, 1914 (superseded combination)
Subgenus Prionospio (Prionospio) [auctorum] accepted as Prionospio Malmgren, 1867 (superseded combination)
  » Species Prionospio (Prionospio) jubata Blake, 1996
  » Species Prionospio (Prionospio) lanceolata Hartmann-Schröder, 1979
  » Species Prionospio (Prionospio) lobulata Fauchald, 1972
  » Species Prionospio (Prionospio) marsupialia Blake, 1996
  » Species Prionospio (Prionospio) orensanzi Blake, 1983
  » Species Prionospio (Prionospio) pacifica Zhou & Li, 2009
  » Species Prionospio (Prionospio) dubia Day, 1961 represented as Prionospio dubia Day, 1961 (disused subgenus)
  » Species Prionospio (Prionospio) anuncata Fauchald, 1972 accepted as Prionospio anuncata Fauchald, 1972 (superseded subgenus)
  » Species Prionospio (Prionospio) australiensis Blake & Kudenov, 1978 accepted as Prionospio australiensis Blake & Kudenov, 1978 (superseded subgenus)
  » Species Prionospio (Prionospio) cristata Foster, 1971 accepted as Prionospio cristata Foster, 1971 (superseded subgenus)
  » Species Prionospio (Prionospio) depauperata Imajima, 1990 accepted as Prionospio depauperata Imajima, 1990 (superseded subgenus)
  » Species Prionospio (Prionospio) ehlersi Fauvel, 1928 accepted as Prionospio ehlersi Fauvel, 1928 (superseded subgenus)
  » Species Prionospio (Prionospio) ergeni Dagli & Çinar, 2009 accepted as Prionospio ergeni Dagli & Çinar, 2009
  » Species Prionospio (Prionospio) heterobranchia Moore, 1907 accepted as Prionospio heterobranchia Moore, 1907 (superseded subgenus)
  » Species Prionospio (Prionospio) lineata Imajima, 1990 accepted as Prionospio lineata Imajima, 1990 (superseded subgenus)
  » Species Prionospio (Prionospio) membranacea Imajima, 1990 accepted as Prionospio membranacea Imajima, 1990 (superseded subgenus)
  » Species Prionospio (Prionospio) oshimensis Imajima, 1990 accepted as Prionospio oshimensis Imajima, 1990 (superseded subgenus)
  » Species Prionospio (Prionospio) paradisea Imajima, 1990 accepted as Prionospio paradisea Imajima, 1990 (superseded subgenus)
  » Species Prionospio (Prionospio) paucipinnulata Blake & Kudenov, 1978 accepted as Prionospio paucipinnulata Blake & Kudenov, 1978
  » Species Prionospio (Prionospio) queenslandica Blake & Kudenov, 1978 accepted as Prionospio queenslandica Blake & Kudenov, 1978 (superseded subgenus)
  » Species Prionospio (Prionospio) steenstrupi Malmgren, 1867 accepted as Prionospio steenstrupi Malmgren, 1867 (superseded subgenus)
  » Species Prionospio (Prionospio) tridentata Blake & Kudenov, 1978 accepted as Prionospio tridentata Blake & Kudenov, 1978 (superseded subgenus)
  » Species Prionospio (Prionospio) tripinnata Maciolek, 1985 accepted as Prionospio tripinnata Maciolek, 1985 (superseded subgenus)
  » Species Prionospio (Prionospio) variegata Imajima, 1990 accepted as Prionospio variegata Imajima, 1990 (superseded subgenus)
Species Prionospio africana Augener, 1918 accepted as Paraprionospio pinnata (Ehlers, 1901) (subjective synonym)
Species Prionospio alata Moore, 1923 accepted as Paraprionospio alata (Moore, 1923) (superseded original combination)
Species Prionospio banyulensis Laubier, 1966 accepted as Aurospio banyulensis (Laubier, 1966) (superseded original combination)
Species Prionospio cirrobranchiata Day, 1961 accepted as Aonidella cirrobranchiata (Day, 1961) (superseded original combination)
Species Prionospio longibranchiata Reish, 1968 accepted as Prionospio delta Hartman, 1965 (subjective synonym)
Species Prionospio multibranchiata Fauvel, 1928 accepted as Prionospio polybranchiata Fauvel, 1929 (replaced primary homonym in Prionospio)
Species Prionospio ockelmanni Pleijel, 1985 accepted as Aurospio banyulensis (Laubier, 1966) (subjective synonym)
Species Prionospio ornata Berkeley & Berkeley, 1961 accepted as Paraprionospio pinnata (Ehlers, 1901) (subjective synonym)
Species Prionospio pilkena Wilson, 1990 accepted as Aurospio pilkena (Wilson, 1990) (superseded original combination)
Species Prionospio pinnata Ehlers, 1901 accepted as Paraprionospio pinnata (Ehlers, 1901) (superseded original combination)
Species Prionospio plumosa Treadwell, 1931 accepted as Prionospio treadwelli Hartman, 1951 accepted as Paraprionospio alata (Moore, 1923) (junior homonym)
Species Prionospio plumosus M. Sars in G.O. Sars, 1872 accepted as Prionospio plumosa M. Sars in G.O. Sars, 1872 (incorrect original spelling for gender agreement)
Species Prionospio salzi Laubier, 1970 accepted as Laubieriellus salzi (Laubier, 1970) (superseded original combination)
Species Prionospio tenuis (Verrill, 1880) accepted as Prionospio steenstrupi Malmgren, 1867 (subsequent combination of subjective synonym)
Species Prionospio treadwelli Hartman, 1951 accepted as Paraprionospio alata (Moore, 1923) (subjective synonym)
marine, brackish, fresh, terrestrial
recent only
Malmgren, A.J. (1867). Annulata Polychaeta Spetsbergiæ, Grœnlandiæ, Islandiæ et Scandinaviæ. Hactenus Cognita. Ex Officina Frenckelliana, Helsingforslæ. 127 pp. & XIV plates., available online at http://www.biodiversitylibrary.org/bibliography/13358
page(s): 93 [details]   
Note Type species P. steenstrupi Malmgren 1867 in...  
From other sources
Type species Type species P. steenstrupi Malmgren 1867 in Fauchald, 1977<246>. [details]
Etymology Prionospio is composed by the Greek noun prion, meaning 'saw', followed by the name of the genus Spio Fabricius, 1785, type...  
Etymology Prionospio is composed by the Greek noun prion, meaning 'saw', followed by the name of the genus Spio Fabricius, 1785, type of the family Spionidae, and refers to the saw-like appearance in profile of the pinnate branchiae of the genotype, Prionospio steenstrupi Malmgren, 1867. [details]
Read, G.; Fauchald, K. (Ed.) (2018). World Polychaeta database. Prionospio Malmgren, 1867. Accessed at: http://www.marinespecies.org/polychaeta/aphia.php?p=taxdetails&id=129620 on 2019-06-17
Date
action
by
2004-12-21 15:54:05Z
created
2008-03-26 11:36:43Z
changed
2016-03-17 02:13:06Z
changed
2016-05-29 12:39:25Z
changed

original description Malmgren, A.J. (1867). Annulata Polychaeta Spetsbergiæ, Grœnlandiæ, Islandiæ et Scandinaviæ. Hactenus Cognita. Ex Officina Frenckelliana, Helsingforslæ. 127 pp. & XIV plates., available online at http://www.biodiversitylibrary.org/bibliography/13358
page(s): 93 [details]   

original description  (of Apoprionospio Foster, 1969) Foster, N.M. (1969). New species of spionids (Polychaeta) from the Gulf of Mexico and Caribbean Sea with a partial revision of the genus <i>Prionospio</i>. <em>Proceedings of the Biological Society of Washington.</em> 82(38): 381-400., available online at http://www.biodiversitylibrary.org/item/107580#page/417/mode/1up
page(s): 383 [details]   

original description  (of Anaspio Chamberlin, 1920) Chamberlin, Ralph V. (1920). The polychaetes collected by the Canadian Arctic Expedition, 1913-18. <em>Report of the Canadian Arctic Expedition 1913-18.</em> 9B: 1-41, plates 1-6., available online at https://biodiversitylibrary.org/page/37673125
page(s): 18 [details]  OpenAccess publication 

original description  (of Aquilaspio Foster, 1971) Foster, Nancy Marie. (1971). Spionidae (Polychaeta) of the Gulf of Mexico and the Caribbean Sea. <em>Studies on the Fauna of Curaçao and other Caribbean Islands.</em> 36(129): 1-183., available online at http://www.repository.naturalis.nl/record/506046
page(s): 105 [details]  Available for editors  PDF available [request] 

original description  (of Periptyches Grube, 1873) Grube, Adolf-Eduard. (1873). Über ein paar neue Anneliden aus der Familie der Spiodeen. <em>Jahres-Bericht der Schlesichen Gesellschaft für vaterländische Cultur.</em> 50: 57-58., available online at https://biodiversitylibrary.org/page/38491131 [details]  OpenAccess publication 

original description  (of Kinbergella McIntosh, 1909) McIntosh, W.C. 1909. Notes from the Gatty Marine Laboratory, St. Andrews. No. 31. 1. On a young stage of Gadus luscus with bold transverse bars of pigment. 2. On the british Spionidae. 3. On the Spionidae dredged by H.M.S. 'Porcupine' in 1869 and 1870. Annals and Magazine of Natural History 3: 153-180. [details]   

taxonomy source Paterson, Gordon L. J.; Neal, Lenka; Altamira, Iris; Soto, Eulogio H.; Smith, Craig R.; Menot, Lenaick; Billett, David S. M.; Cunha, Marina R.; Marchais-Laguionie, Clare; Glover, Adrian G. (2016). New <em>Prionospio</em> and <em>Aurospio</em> Species from the Deep Sea (Annelida: Polychaeta). <em>Zootaxa.</em> 4092(1): 1-32., available online at https://doi.org/10.11646/zootaxa.4092.1.1 [details]   

basis of record Bellan, G. (2001). Polychaeta, <i>in</i>: Costello, M.J. <i>et al.</i> (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. <em>Collection Patrimoines Naturels.</em> 50: 214-231. (look up in IMIS[details]   

additional source Day, J. H. (1967). [Sedentaria] A monograph on the Polychaeta of Southern Africa. Part 2. Sedentaria. British Museum (Natural History), London. pp. 459–842., available online at http://www.biodiversitylibrary.org/bibliography/8596  [details]   

additional source Sigvaldadóttir, E. (1998). Cladistic analysis and classification of <i>Prionospio</i> and related general (Polychaeta, Spionidae). <em>Zoologica Scripta.</em> 27(3): 175-187., available online at http://dx.doi.org/10.1111/j.1463-6409.1998.tb00435.x [details]   

additional source Glasby, C.J.; Read, G.B.; Lee, K.E.; Blakemore, R.J.; Fraser, P.M.; Pinder, A.M.; Erséus, C.; Moser, W.E.; Burreson, E.M.; Govedich, F.R.; Davies, R.W.; Dawson, E.W. (2009). Phylum Annelida: bristleworms, earthworms, leeches, in: Gordon, D.P. (Ed.) (2009). New Zealand inventory of biodiversity: 1. Kingdom Animalia: Radiata, Lophotrochozoa, Deuterostomia. pp. 312-358. [details]   

additional source Fauchald, K. (1977). The polychaete worms, definitions and keys to the orders, families and genera. <em>Natural History Museum of Los Angeles County: Los Angeles, CA (USA), Science Series.</em> 28:1-188., available online at http://www.vliz.be/imisdocs/publications/123110.pdf [details]   

identification resource Delgado-Blas, V. H. 2014. Redescriptions and reestablishments of some species belonging to the genus Prionospio (Polychaeta, Spionidae) and descriptions of three new species. Helgoland Marine Research 68(1): 113-132, available online at https://doi.org/10.1007/s10152-013-0372-1
page(s): [Key to known species of Prionospio with five pairs of branchiae] [details]   

identification resource Delgado-Blas, Víctor Hugo. (2015). <em>Prionospio</em> (Polychaeta, Spionidae) from the Grand Caribbean Region, with the descriptions of five new species and a key to species recorded in the area. <em>Zootaxa.</em> 3905(1): 69-90., available online at https://doi.org/10.11646/zootaxa.3905.1.4
note: Caribbean species key [details]   

emendation (re-diagnosis of genus) Peixoto, Antônio João Malafaia; Paiva, Paulo Cesar de. (2019). New <i>Prionospio</i> and <i>Laubieriellus</i> (Annelida: Spionidae) species from Southeastern Brazil. <em>Zootaxa.</em> 4577(3): 529-547., available online at https://doi.org/10.11646/zootaxa.4577.3.7
page(s): 531-532; note: Genus diagnosis emended to include "Branchiae from chaetiger 2, chaetiger 3 or rarely absent. Branchiae limited to anterior chaetigers, 0-15 pairs [...]." [details]  Available for editors  PDF available [request] 
 

From editor or global species database
Diagnosis Original diagnosis by Malmgren (1867: 93): ''Corpus elongatum sublineare postice sensim paullum attenuatam. Lobus cephalicus brevis parte frontale parum elevata subrectangulare apice transversa. Antennae nullae. Tentaculum haud conspicuum. Oculi nulli. Tubercula setigera similia pinnis compressis parum discretis, labiis minutis, magnis foliaceis modo in segmentis 4 anticis obviis. Branchiae, paria 4, liberae in segmentis 4 anticis obviae, dorso adnatae, paris primi et quarti aequales ceteris longiores pinnatae, paris secundi et tertii multo breviores aequales filiformes. Setae in segmentis anticis modo capillares in utraque pinna, in posticis segmentis capillares et uncinatae in pinna ventrali, modo capillares in dorsali. Setae capillares versus apicem attenuatum curvatum anguste limbatae in segmentis anticis, in posticis rectae tenuissime attenuatae haud limbatae. Uncini rostrati apice limbati, vertice unidentati. Seta infima rami inferioris in segmentis mediis linearis falcis instar curvata.''
 [details]

Etymology Prionospio is composed by the Greek noun prion, meaning 'saw', followed by the name of the genus Spio Fabricius, 1785, type of the family Spionidae, and refers to the saw-like appearance in profile of the pinnate branchiae of the genotype, Prionospio steenstrupi Malmgren, 1867. [details]

From other sources
Habitat Known from seamounts and knolls [details]

Type species Type species P. steenstrupi Malmgren 1867 in Fauchald, 1977<246>. [details]