Traits name details
original description
Nuttall, C. P. (1990). A review of the Tertiary non-marine molluscan faunas of the Pebasian and other inland basins of north-western South America. <em>Bulletin of The British Museum (Natural History), Geology Series.</em> 45(2): 165-371., available online at https://www.biodiversitylibrary.org/page/45474563 page(s): 207-208, figs 147-148 [details]
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