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Hayward, P.J. (2001). Bryozoa, in: Costello, M.J. et al. (Ed.) European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 325-333
In: Costello, M.J.; Emblow, C.; White, R.J. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50. Muséum national d'Histoire naturelle: Paris. ISBN 2-85653-538-0. 463 pp., more
In: Collection Patrimoines Naturels. Muséum national d'Histoire naturelle: Paris. ISSN 1158-422X

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    Bryozoa [WoRMS]

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  • Hayward, P.J., more

    This catalogue of European Bryozoa incorporates the checklists prepared by Hayward (1997) and Brattegard (1995, also checked by Hayward), the species lists in Hayward and Ryland (1998, 1999), Zabala and Maluquer (1988), and relevant primary sources published after 1985. All faunistic and systematic papers relating to shelf and coastal European Bryozoa listed in the Zoological Record from 1985 to date have been consulted, but only those including new species or new European records are referenced here. Synonyms of most species included are to be found in the sources cited above; an unsourced entry indicates either a recently described species or unpublished records held by the compiler. The ERMS list was checked by J. Harmelin.

    Taxonomic and faunistic research on European Bryozoa has been continuous through the last three decades, but geographical coverage and frequency of investigation have been extremely variable. The British Isles, Ireland and the Atlantic coasts of north-west Europe have attracted most research effort and the faunas of these regions are reasonably well known, although completely new species continue to be discovered even in such well studied areas as the English Channel (Reverter-Gil and Fernandez-Pulpeiro, 1996; Hayward and Hansen, 1999). The Faeroe Islands have been the subject of two surveys (Kramp, 1934; Hayward, 1994), the latter resulting in 15 new species and 13 new records for the temperate north-east Atlantic. The faunas of Iceland and west Greenland are similarly poorly known but are currently under review. The synoptic key of Zabala and Maluquer (1988) was founded on numerous sources relating to the western Mediterranean and Adriatic faunas, on the few available accounts of Aegean Bryozoa and the even sparser records for the southern coasts of the Mediterranean. Mediterranean Bryozoa are still only incompletely known, at best, and it is certain that many more undescribed species will be discovered as research continues.


    • Alvarez, J. A. 1987. Notas sobre la fauna briozoologica marina Iberica. II. Especies de la Costa Vasca: estudio zoogeographico. KOBIE (Serie Ciencias Naturales) Bilbao, 16: 215 - 222
    • Alvarez, J. A. 1990. Una coleccion de briozoos procedentes de la costa de Alicante. Boletin Inst. Esp. Oceanog., 6: 21 - 40
    • Alvarez, J. A. 1992. Sobre algunas especies de la familia Lichenoporidae Smitt, 1866 (Bryozoa, Cyclostomida) en la region Atlantico-Mediterranea. Parte I: genero Disporella
    • Gray, 1848. Cahiers de Biologie Marine, 33: 201 - 243
    • Alvarez, J. A. 1993, Fenestrulina barrosoi sp. nov. (Bryozoa: Cheilostomida) with a review of the genus Fenestrulina on the Iberian Peninsula. Journal of the Marine Biological Association of the U. K., 73: 831 - 835
    • Alvarez, J. A. 1994. La famille des Lichenoporidae (Bryozoa, Cyclostomida) dans les provinces Atlantico-Mediterraneenne et boreale Partie III: etude des collections du Museum National d’Histoire Naturelle de Paris. Cahiers de Biologie Marine, 35: 491 - 509
    • Alvarez, J. A. 1995. New data on the family Lichenoporidae Smitt (Bryozoa: Cyclostomida) from the Mediterranean region. Journal of Natural History, 29: 1067 - 1079.
    • Aristegui, J. 1984. Briozoos Quilostomados (Ectoprocta, Cheilostomata) de Canarias: estudio sistematiico, faunistico y biogeografico. Tesis doctoral, Universidad de La Laguna. 524 pp.
    • Balduzzi, A., Barbieri, M. and Gristina, M. 1991. Morphology and life strategies of Aetea (Bryozoa: Cheilostomata) living on some western Mediterranean Posidonia oceanica meadows. Bulletin de la Societe des Sciences Naturelles de l’Ouest de la France, Memoire HS 1: 1 - 12.
    • Brattegard, T. 1995.Bryozoa. Pp. 278 - 295 in Brattegard, T. and Holthe, T. (eds), Distribution of marine, benthic macro-organisms in Norway. Research Report for DN 1997-1. Directorate for Nature Management: Trondheim.
    • Fernandez-Pulpeiro, E., Reverter-Gil, O. and Ramil, F. 1990. Inventario de los briozoos de Galicia. Thalassas, 8: 57 - 67
    • Harmelin, J.-G. 1997. Especes affines microsympatriques chez Puellina (Bryozoa, Cheilostomata) et description d’especes nouvelles. Zoologica Scripta, 17: 25 - 38
    • Harmelin, J.-G. and Aristegui, J. 1988. New Cribrilinidae (Bryozoa, Cheilostomata) from the upper bathyal of the Atlanto-Mediterranean region. Journal of Natural History, 22: 507 - 535
    • Harmelin, J.-G. and d’Hondt, J.-L. 1992a. Bryozoaires des parages de Gibraltar (campagne oceanographique BALGIM, 1984) 1 - Cheilostomes.Bulletin du Museum National d’Histoire Naturelle, Paris, serie 4, 14: 23 - 67
    • Harmelin, J.-G. and d’Hondt, J.-l. 1992b. Bryozoaires des parages de Gibraltar (campagne oceanographique Balgim, 1984) 2 - Ctenostomes et Cyclostomes. Bulletin du Museum National d’Histoire Naturelle, Paris, serie 4, 14: 605 - 621
    • Harmelin, J.-G., Boronat, J., Moisette, P. and Rosso, A. 1989. Distansescharella seguenzai Cipolla, 1921 (Bryozoa, Cheilostomata), nouvelles donnees morphologiques et ecologiques tirees de specimens fossiles (Miocene, Pliocene) et actuels de Mediterranee. Geobios, 22: 484 - 501
    • Hayward, P. J. 1994. New species and new records of cheilostomatous Bryozoa from the Faroe Islands, collected by BIOFAR. Sarsia, 79: 181 - 206
    • Hayward, P. J. 1997. Bryozoa. Pp. 273 - 282, in Howson, C. M. and Picton, B. E. (eds), The species directory of the marine fauna and flora of the British Isles and surrounding seas. Belfast and Ross-on-Wye: Ulster Museum and the Marine Conservation Society.
    • Hayward, P. J. and Hansen, K. B. 1999. Three newly recognised cheilostomate bryozoans from the British sea area. Journal of the Marine Biological Association of the U.K. 79: 917 - 921
    • Hayward, P. J. and Ryland, J. S. 1998. Cheilostomatous Bryozoa. Part 1, Aeteoidea - Cribrilinoidea. Synopses of the British Fauna (new series), 10 (second edition): 1 - 366
    • Hayward, P. J. and Ryland, J. S. 1999. Cheilostomatous Bryozoa. Part 2, Hippothooidea - Celleporoidea. Synopses of the British Fauna (new series), 14 (second edition): 1 - 416
    • Hondt, J.-L. d’ 1984. Un nouvel immigrant dans le Bassin d’Arcachon, Watersipora aterrima (Ortmann, 1890) (Bryozoaire Cheilostome). Congres national des Societes savantes, Dijon, 1984, sciences, fasc. II: 237 - 245
    • Hondt, J.-L. d’ 1988. Bryozoa from the coast of Israel. Boll. Zool. 3: 191 - 203Kramp, P. L. 1934. Bryozoa. Zoology of the Faroes, 58: 1 - 39
    • Reverter-Gil, O. 1994. Briozoos de la Ria de Ferrol. Tesis doctoral, Universidad de Santiago de Compostela.
    • Reverter-Gil, O. and Fernandez-Pulpeiro, E. 1996. Cribrilinidae from the Ria de Ferrol (NW Spain). Journal of Natural History, 30: 1247 -1260
    • Reverter-Gil, O. and Fernandez-Pulpeiro, E. 1997. Two new species of Schizomavella (Bryozoa, Cheilostomatida). Cahiers de Biologie Marine, 38: 1 - 6
    • Reverter- Gil, O., d’Hondt, J.-L. and Fernandez-Pulpeiro, E. 1995. Mise a jour de l’inventaire des bryozoaires de Roscoff publie par Echalier et Prenant (1951). Cahiers de Biologie Marine, 36: 123 - 131
    • Rosso, A. 1996. Popolamenti e Tanatocenosi a briozoi di fondi mobili circalitorali del Golfo di Noto (Sicilia, Italia). Naturalista sicil. S. IV, 30: 189 - 225
    • Rosso, A. 1999. Recent and fossil species of Characodoma Maplestone, 1900 (Bryozoa) from the Mediterranean, with description of two new species. Journal of Natural History, 33: 415 - 437
    • Zabala, M. and Maluquer, P. 1988. Illustrated keys for the classification of Mediterranean Bryozoa. Treballs del Museu de Zoologia, Barcelona, 4: 1 - 294
    • Zabala, M., Maluquer, P. and Harmelin, J.-G. 1993. Epibiotic bryozoans on deep-water scleractinian corals from the Catalonia slope (western Mediterranean, Spain, France). Scientia Marina, 57: 65 - 78

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