Traits name details
original description
Brown, A. P. & Pilsbry, H. A. (1913). Fauna of the Gatun Formation, Isthmus of Panama - II. <em>Proceedings of the Academy of Natural Sciences of Philadelphia.</em> 64: 500–519, pls. 22–26., available online at https://www.biodiversitylibrary.org/page/1695158 page(s): 506, pl. 22, fig. 5 [details] 
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