Traits name details
original description
Horst, R. (1893). Descriptions of earthworms. <em>Notes from the Leyden Museum.</em> 15(4): 316-329 + Plate 10., available online at https://www.biodiversitylibrary.org/page/12957429 page(s): 327 [details] 
basis of record
Reynolds, J. W.; Wetzel, M. J. (2018). Nomenclatura Oligochaetologica – A catalogue of names, descriptions and type specimens. Editio Secunda. , available online at https://nomenclatura-oligochaetologica.inhs.illinois.edu/ [details]
additional source
Beddard, F.E. (1895). A monograph of the order of Oligochaeta. <em>[Book].</em> 769 pp. The Clarendon Press, Oxford., available online at https://www.biodiversitylibrary.org/item/71343 page(s): 433 [details]
new combination reference
Michaelsen, W. (1916). Oligochäten aus dem Naturhistorischen Reichsmuseum zu Stockholm. <em>Arkiv för zoologi.</em> 10(9): 1-21., available online at https://www.biodiversitylibrary.org/page/6411988 page(s): 13 [details]
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