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Erpenbeck, D.; Sutcliffe, P.; Cook, S. de C.; Dietzel, A.; Maldonado, M.; Van Soest, R.W.M.; Hooper, J.N.A.; Wörheide, G. (2012). Horny sponges and their affairs: On the phylogenetic relationships of keratose sponges. Molecular Phylogenetics and Evolution. 63 (3): 809-816.