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Longshaw, M., Pursglove, M. & Shinn, A.P. (2003). Gyrodactylus quadratidigitus n. sp. (Monogenea: Gyrodactylidae), a parasite of the leopard-spotted goby Thorogobius ephippiatus (Lowe) from the south-western coast of the UK. Journal of Natural History 55, 151-157.