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Ohtsuka, S. & K. Kimoto. (1989). Tortanus (Atortus) (Copepoda: Calanoida) of southern Japanese waters, with description of two new species, T. (A.) digitalis and T. (A.) ryukyuensis, and discussion on distribution and swarming behavior of Atortus. Journal of Crustacean Biology 9(3):392-408.