Intertidal. [details]
Known only from the type locality. [details]
Intertidal gravel. [details]
Database data entry spelling error as 'pustulatus' corrected February 2019. [details]
Pacific Ocean, Peru, Lima (geocoordinates not provided, but estimated with gazetteer to be approximately lat. ... [details]
El Salvador. Several localities are given, but a type locality is not stated. It might be determined from the ... [details]