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Díaz-Tapia, P.; McIvor, L.; Freshwater, D. W.; Verbruggen, H.; Wynne, M. J.; Maggs, C. A. (2017). The genera Melanothamnus Bornet & Falkenberg and Vertebrata S.F. Gray constitute well-defined clades of the red algal tribe Polysiphonieae (Rhodomelaceae, Ceramiales). European Journal of Phycology. 52(1): 1-30.