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Cardoso, Y. P.; Almirón, A.; Casciotta, J.; Aichino, D.; Lizarralde, M. S.; Montoya-Burgos, J. I. (2012). Origin of species diversity in the catfish genus Hypostomus (Siluriformes: Loricariidae) inhabiting the Paraná river basin, with the description of a new species. Zootaxa. 3453: 69-83.