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Bührnheim, C. M.; Carvalho, T. P.; Malabarba, L. R.; Weitzman, S. H. (2008). A new genus and species of characid fish from the Amazon basin: the recognition of a relictual lineage of characid fishes (Ostariophysi: Cheirodontinae: Cheirodontini). Neotropical Ichthyology. 6(4): 663-678.