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Antal, L.; László, B.; Kotlík, P.; Mozsár, A.; Czeglédi, I.; Oldal, M.; Kemenesi, G.; Jakab, F.; Nagy, S. A. (2016). Phylogenetic evidence for a new species of Barbus in the Danube River basin. Molecular Phylogenetics and Evolution. 96: 187-194.