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Shih, H.-T.; Ng, P. K. L.; Davie, P. J. F.; Schubart, C. D.; Türkay, M.; Naderloo, R.; Jones, D. S.; Liu, M.-Y. (2016). Systematics of the family Ocypodidae Rafinesque, 1815 (Crustacea: Brachyura), based on phylogenetic relationships, with a reorganization of subfamily rankings and a review of the taxonomic status of Uca Leach, 1814, sensu lato and its subgenera. Raffles Bulletin of Zoology, (Raffles Bull. Zool.). 64: 139-175.