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Brandão, S. N.; Yasuhara, M.; Irizuki, T.; Horne, D. J. (2013). The ostracod genus Trachyleberis (Crustacea; Ostracoda) and its type species. Marine Biodiversity. 43: 363-405.
170403
Brandão, S. N.; Yasuhara, M.; Irizuki, T.; Horne, D. J.
2013
The ostracod genus Trachyleberis (Crustacea; Ostracoda) and its type species
Marine Biodiversity
43: 363-405
Publication
Available for editors  PDF available [request]
The Trachyleberididae is one of the most diverse families of the Ostracoda in Mesozoic to present-day marine environments. Its type genus is Trachyleberis Brady, 1898, first described on the basis of specimens collected alive. Unfortunately the identity and morphological characteristics of the type species of Trachyleberis have long been the subject of confusion resulting from misidentifications and misunderstandings in the literature. We review the taxonomic history of this problem, providing extensive synonymies, new diagnoses, descriptions and illustrations of relevant species. We establish the identity of the type species of Trachyleberis, Cythere scabrocuneata Brady, 1880, by reference to a previously-designated lectotype from the Seto Inland Sea, Japan, noting that a second distinct species, Trachyleberis niitsumai Ishizaki, 1971, is represented in the original syntypic series. We consider that in the interests of nomenclatural and taxonomic stability C. scabrocuneata should be retained as the type species of Trachyleberis, despite the fact that the original description of the genus was based on a species misidentified as C. scabrocuneata and which was subsequently described as a new species, Trachyleberis lytteltonensis Harding and Sylvester-Bradley, 1953. The geographical and stratigraphical distributions of Trachyleberis are reconsidered in the light of our revision. Trachyleberis now comprises 18 known species inhabiting shallow marine environments of the Northwestern Pacific from Japan to the northern coast of the South China Sea off Hong Kong. The revised stratigraphic occurrence of Trachyleberis is from the Eocene to Recent. As many as 400 living and fossil species have been previously assigned to Trachyleberis and they range from the Cretaceous to the Recent; Cretaceous species formerly attributed to Trachyleberis are now reassigned to other genera. We suggest that Trachyleberis most likely evolved from a Cythereis-like ancestor during the Eocene–Oligocene of Japan, the same area that is the centre of its modern distribution.
Biodiversity, Taxonomic and ecological diversity
Paleontology, Fossils, Paleobiology
Systematics, Taxonomy
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2013-10-20 19:45:18Z
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2016-07-20 09:03:18Z
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Costa edwardsii edwardsii (Roemer, 1838) Keij, 1957 † (additional source)
Costa edwardsii padana (Ruggieri, 1950) † (additional source)
Cythere echinata elongata Lienenklaus, 1894 † accepted as Trachyleberis echinata elongata (Lienenklaus, 1894) † (additional source)
Cythere scabrocuneata Brady, 1880 (new combination reference)
Rectotrachyleberis edwardsii (Roemer, 1838) † accepted as Costa edwardsii (Roemer, 1838) † (additional source)
Rectotrachyleberis edwardsii edwardsii (Roemer, 1838) † accepted as Costa edwardsii edwardsii (Roemer, 1838) Keij, 1957 † (additional source)
Rectotrachyleberis elegantissima (Lienenklaus, 1894) Stancheva, 1962 † (additional source)
Trachyleberididae Sylvester-Bradley, 1948 (subsequent type designation)
Trachyleberis Brady, 1898 (subsequent type designation)
Trachyleberis (Acanthocythereis) Howe, 1963 † accepted as Acanthocythereis Howe, 1963 (additional source)
Trachyleberis (Acanthocythereis) araneosa (Howe, 1963) Siddiqui, 1971 † accepted as  † (additional source)
Trachyleberis (Acanthocythereis) decoris Siddiqui, 1971 † accepted as  † (additional source)
Trachyleberis (Acanthocythereis) pedigaster Siddiqui, 1971 † (additional source)
Trachyleberis (Acanthocythereis) postcornis Siddiqui, 1971 † accepted as  † (additional source)
Trachyleberis (Acanthocythereis) procapsus Siddiqui, 1971 † accepted as  † (additional source)
Trachyleberis (Acanthocythereis) usitata Siddiqui, 1971 † (additional source)
Trachyleberis (Actinocythereis) Puri, 1953 accepted as Actinocythereis Puri, 1953 (additional source)
Trachyleberis (Actinocythereis) birmanica Gramann, 1975 † accepted as Actinocythereis birmanica (Gramann, 1975) † (additional source)
Trachyleberis (Actinocythereis) birmanica birmanica Gramann, 1975 † accepted as Actinocythereis birmanica birmanica (Gramann, 1975) † (additional source)
Trachyleberis (Actinocythereis) birmanica kyaukkokensis Gramann, 1975 † accepted as Actinocythereis birmanica kyaukkokensis (Gramann, 1975) † (additional source)
Trachyleberis (Actinocythereis) birmanica pyawbwe Gramann, 1975 † accepted as Actinocythereis birmanica pyawbwe (Gramann, 1975) † (additional source)
Trachyleberis (Actinocythereis) exanthemata (Ulrich & Bassler, 1904) † accepted as Actinocythereis exanthemata (Ulrich & Bassler, 1904) Puri, 1953 † (additional source)
Trachyleberis (Actinocythereis) marchilensis Hartmann, 1965 (additional source)
Trachyleberis (Actinocythereis) scutigera (Brady, 1868) Gramann, 1975 accepted as Actinocythereis scutigera (Brady, 1868) (additional source)
Trachyleberis (Actinocythereis) tumefacentis (Luebimova & Guha, 1960) Krstic, 1979 † (additional source)
Trachyleberis (Actinocythereis) wechesensis (Sutton & Williams, 1940) † accepted as Actinocythereis wechesensis (Sutton & Williams, 1940) Puri, 1953 † (additional source)
Trachyleberis (Actinocythereis) woodwardsensis (Howe & Law, 1936) † accepted as Actinocythereis woodwardsensis (Howe & Law, 1936) Puri, 1953 † (additional source)
Trachyleberis (Alteratrachyleberis) Li, 1964 accepted as Alteratrachyleberis Li, 1964 † (new combination reference)
Trachyleberis (Alteratrachyleberis) quasiperforata Kulichenko, 1976 † (additional source)
Trachyleberis (Costa) Neviani, 1928 † accepted as Costa Neviani, 1928 † (additional source)
Trachyleberis (Costa) costata Scheremeta, 1969 † (additional source)
Trachyleberis (Costa) edwardsii (Roemer, 1838) Keij, 1957 † accepted as Costa edwardsii (Roemer, 1838) † (additional source)
Trachyleberis (Costa) elongata Scheremeta, 1969 † accepted as  † (additional source)
Trachyleberis (Echinocythereis) Puri, 1954 † accepted as Echinocythereis Puri, 1954 (additional source)
Trachyleberis (Echinocythereis) scabra (Muenster, 1830) Li, 1966 accepted as Echinocythereis scabra (Muenster, 1830) (additional source)
Trachyleberis (Henryhowella) Puri, 1957 accepted as Henryhowella Puri, 1957 (additional source)
Trachyleberis (Hermanites) Puri, 1955 accepted as Hermanites Puri, 1955 (additional source)
Trachyleberis (Hermanites) bassleri (Ulrich, 1901) Li, 1966 † accepted as  † (additional source)
Trachyleberis (Hermanites) scribrosa (Li, 1963) Li, 1966 † accepted as  † (additional source)
Trachyleberis (Hermanites) suzakensis daedalia (Li, 1963) Li, 1966 † (additional source)
Trachyleberis (Hermanites) suzakensis suzakensis (Mandelstam, 1959) Li, 1966 † (additional source)
Trachyleberis (Hermanites) torosa (Li, 1963) Li, 1966 † accepted as  † (additional source)
Trachyleberis (Rectotrachyleberis) Ruggieri, 1952 accepted as Rectotrachyleberis Ruggieri, 1952 (new combination reference)
Trachyleberis (Rectotrachyleberis) edwardsii (Roemer, 1838) Ruggieri, 1952 † accepted as Costa edwardsii (Roemer, 1838) † (additional source)
Trachyleberis (Rectotrachyleberis) edwardsii edwardsii (Roemer, 1838) Ruggieri, 1952 † accepted as Costa edwardsii edwardsii (Roemer, 1838) Keij, 1957 † (additional source)
Trachyleberis (Rectotrachyleberis) edwardsii padana (Ruggieri, 1950) Ruggieri, 1952 accepted as Costa edwardsii padana (Ruggieri, 1950) † (additional source)
Trachyleberis (Rectotrachyleberis) hamata (Mueller, 1894) Ruggieri, 1952 accepted as Carinocythereis hamata (Mueller, 1894) (additional source)
Trachyleberis (Rectotrachyleberis) polytrema (Brady, 1878) Ruggieri, 1952 accepted as Costa polytrema (Brady, 1878) Ruggieri, 1959 (additional source)
Trachyleberis (Rectotrachyleberis) tricostata (Reuss, 1850) Ruggieri, 1952 † accepted as Costa (Cuneocosta) tricostata (Reuss, 1850) Ruggieri, 1992 † (additional source)
Trachyleberis (Trachyleberidea) Bowen, 1953 accepted as Trachyleberidea Bowen, 1953 † (additional source)
Trachyleberis (Trachyleberis) Brady, 1898 (new combination reference)
Trachyleberis (Trachyleberis) aculeata (Bosquet, 1852) Apostolescu, 1955 † accepted as  † (additional source)
Trachyleberis (Trachyleberis) arachnoidea (Bosquet, 1852) Keij, 1957 † represented as Trachyleberis arachnoidea (Bosquet, 1852) Keij, 1957 † (additional source)
Trachyleberis (Trachyleberis) asperrima (Reuss, 1850) Keij, 1957 † accepted as Henryhowella asperrima (Reuss, 1850) Bold, 1960 † (additional source)
Trachyleberis (Trachyleberis) asperrima asperrima (Reuss, 1850) Keij, 1957 † accepted as Henryhowella asperrima asperrima (Reuss, 1850) Bold, 1960 † (additional source)
Trachyleberis (Trachyleberis) asperrima echinata (Reuss, 1851) Keij, 1957 † accepted as  † (additional source)
Trachyleberis (Trachyleberis) bimammillata Siddiqui, 1971 † represented as Trachyleberis bimammillata Siddiqui, 1971 † (additional source)
Trachyleberis (Trachyleberis) dunelmensis (Norman, 1865) accepted as Actinocythereis dunelmensis (Norman, 1865) (additional source)
Trachyleberis (Trachyleberis) echinata (Reuss, 1851) Keij, 1957 † represented as  † (additional source)
Trachyleberis (Trachyleberis) hystrix (Reuss, 1850) Ruggieri, 1950 † (additional source)
Trachyleberis (Trachyleberis) lobuculus Siddiqui, 1971 † represented as Trachyleberis lobuculus Siddiqui, 1971 † (additional source)
Trachyleberis (Trachyleberis) lyelliana (Bosquet, 1852) Keij, 1957 † accepted as  † (additional source)
Trachyleberis (Trachyleberis) scabrocuneata (Brady, 1880) Brady, 1898 represented as Trachyleberis scabrocuneata (Brady, 1880) Brady, 1898 (new combination reference)
Trachyleberis (Trachyleberis) spinosa (Lienenklaus, 1900) Keij, 1957 † accepted as Acanthocythereis spinosa (Lienenklaus, 1900) Moos, 1968 † (additional source)
Trachyleberis acanthoderma (Brady, 1880) Ruan & Hao (Yi-Chun), 1988 accepted as Legitimocythere acanthoderma (Brady, 1880) (additional source)
Trachyleberis aculeata Rosyjeva, 1962 † (additional source)
Trachyleberis aculeata (Bosquet, 1852) Apostolescu, 1955 † accepted as  † (additional source)
Trachyleberis aculeata (Bosquet, 1852) Deroo, 1966 † accepted as  † (additional source)
Trachyleberis aculeata aculeata (Bosquet, 1852) Apostolescu, 1955 † accepted as  † (additional source)
Trachyleberis aculeata horridula (Bosquet, 1854) Margerie, 1968 † (additional source)
Trachyleberis aculeata minituberculata Margerie, 1968 † (additional source)
Trachyleberis aculeata modesta Van Hinte, 1962 † (additional source)
Trachyleberis acuminata Holden, 1964 † (additional source)
Trachyleberis adornata Chochlova, 1968 † accepted as  † (additional source)
Trachyleberis anteplana Bate, 1972 † (additional source)
Trachyleberis arachnoidea (Bosquet, 1852) Keij, 1957 † (additional source)
Trachyleberis artschmanensis Rosyjeva, 1962 † (additional source)
Trachyleberis ascolii Puri, 1963 accepted as Acanthocythereis ascolii (Puri, 1963) † (additional source)
Trachyleberis asperrima (Reuss, 1850) Keij, 1957 † accepted as Henryhowella asperrima (Reuss, 1850) Bold, 1960 † (additional source)
Trachyleberis asperrima asperrima (Reuss, 1850) Keij, 1957 † accepted as Henryhowella asperrima asperrima (Reuss, 1850) Bold, 1960 † (additional source)
Trachyleberis asperrima echinata (Reuss, 1851) Keij, 1957 † accepted as  † (additional source)
Trachyleberis avitis (Schneider, 1953) Uljanova, 1965 † (additional source)
Trachyleberis babadurmasensis Rosyjeva, 1962 † (additional source)
Trachyleberis bahamensis (Brady, 1870) Puri, 1953 (additional source)
Trachyleberis barbata Gramm, 1963 † (additional source)
Trachyleberis barbatus Gramm, 1963 † accepted as Trachyleberis barbata Gramm, 1963 † (additional source)
Trachyleberis baruni Banerji, 1970 † (additional source)
Trachyleberis bassleri (Ulrich, 1901) Brown, 1958 † accepted as  † (additional source)
Trachyleberis bassleri (Ulrich, 1901) Swain, 1952 † accepted as  † (additional source)
Trachyleberis bathymarina Ayress, 1993 accepted as Ayressoleberis bathymarina (Ayress, 1993) (additional source)
Trachyleberis baturini Schneider, 1959 † (additional source)
Trachyleberis bermudezi (Bold, 1946) Bold, 1960 † accepted as Legitimocythere bermudezi (Bold, 1946) † (additional source)
Trachyleberis bermudezi bermudezi (Bold, 1946) Bold, 1960 † accepted as Legitimocythere bermudezi bermudezi (Bold, 1946) † (additional source)
Trachyleberis bermudezi crebripustulosa Bold, 1966 † accepted as Legitimocythere bermudezi crebripustulosa (Bold, 1966) (additional source)
Trachyleberis bhujensis Tewari & Tandon, 1960 † accepted as  † (additional source)
Trachyleberis bicarinata (Swain, 1946) † accepted as Quadracythere bicarinata (Swain, 1946) † accepted as Jugosocythereis bicarinata (Swain, 1946) † (additional source)
Trachyleberis bimammillata Siddiqui, 1971 † (additional source)
Trachyleberis bodjonegoroensis (Kingma, 1948) † (additional source)
Trachyleberis bollii Bold, 1960 † accepted as Bathypterocythereis bollii (Bold, 1960) Coles & Whatley, 1989 † (additional source)
Trachyleberis bowerbankiana (Jones, 1857) Bowen, 1953 † accepted as Agrenocythere bowerbankiana (Jones, 1857) King, 1981 † (additional source)