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NameBrandão, S.N., Yasuhara, M., Irizuki, T. and Horne, D.J. (2013a) The ostracod genus Trachyleberis (Crustacea; Ostracoda) and its type species. Marine Biodiversity 43, 363–405.
AuthorsBrandão, S. N.; Yasuhara, M.; Irizuki, T.; Horne, D. J.
Year2013
TitleThe ostracod genus Trachyleberis (Crustacea; Ostracoda) and its type species
JournalMarine Biodiversity
Suffix43: 363-405
Type Publication
Linkhttp://link.springer.com/10.1007/s12526-013-0163-6
Full textAvailable for editors  PDF available [request]
AbstractThe Trachyleberididae is one of the most diverse families of the Ostracoda in Mesozoic to present-day marine environments. Its type genus is Trachyleberis Brady, 1898, first described on the basis of specimens collected alive. Unfortunately the identity and morphological characteristics of the type species of Trachyleberis have long been the subject of confusion resulting from misidentifications and misunderstandings in the literature. We review the taxonomic history of this problem, providing extensive synonymies, new diagnoses, descriptions and illustrations of relevant species. We establish the identity of the type species of Trachyleberis, Cythere scabrocuneata Brady, 1880, by reference to a previously-designated lectotype from the Seto Inland Sea, Japan, noting that a second distinct species, Trachyleberis niitsumai Ishizaki, 1971, is represented in the original syntypic series. We consider that in the interests of nomenclatural and taxonomic stability C. scabrocuneata should be retained as the type species of Trachyleberis, despite the fact that the original description of the genus was based on a species misidentified as C. scabrocuneata and which was subsequently described as a new species, Trachyleberis lytteltonensis Harding and Sylvester-Bradley, 1953. The geographical and stratigraphical distributions of Trachyleberis are reconsidered in the light of our revision. Trachyleberis now comprises 18 known species inhabiting shallow marine environments of the Northwestern Pacific from Japan to the northern coast of the South China Sea off Hong Kong. The revised stratigraphic occurrence of Trachyleberis is from the Eocene to Recent. As many as 400 living and fossil species have been previously assigned to Trachyleberis and they range from the Cretaceous to the Recent; Cretaceous species formerly attributed to Trachyleberis are now reassigned to other genera. We suggest that Trachyleberis most likely evolved from a Cythereis-like ancestor during the Eocene–Oligocene of Japan, the same area that is the centre of its modern distribution.
Bibliographical
terms
Biodiversity, Taxonomic and ecological diversity
Paleontology, Fossils, Paleobiology
Systematics, Taxonomy
Taxonomic
term
Ostracoda
Taxa
(384)
Checked: verified by a taxonomic editorCosta edwardsii edwardsii (Roemer, 1838) Keij, 1957 † (additional source)
Checked: verified by a taxonomic editorCosta edwardsii padana (Ruggieri, 1950) † (additional source)
Checked: verified by a taxonomic editorCythere echinata elongata Lienenklaus, 1894 † accepted as Checked: verified by a taxonomic editorTrachyleberis echinata elongata (Lienenklaus, 1894) † (additional source)
Checked: verified by a taxonomic editorCythere scabrocuneata Brady, 1880 (new combination reference)
Checked: verified by a taxonomic editorRectotrachyleberis edwardsii (Roemer, 1838) Oertli, 1956 † accepted as Checked: verified by a taxonomic editorCosta edwardsii (Roemer, 1838) † (additional source)
Checked: verified by a taxonomic editorRectotrachyleberis edwardsii edwardsii (Roemer, 1838) Oertli, 1956 † accepted as Checked: verified by a taxonomic editorCosta edwardsii edwardsii (Roemer, 1838) Keij, 1957 † (additional source)
Checked: verified by a taxonomic editorRectotrachyleberis elegantissima (Lienenklaus, 1894) Stancheva, 1962 † (additional source)
Checked: verified by a taxonomic editorTrachyleberididae Sylvester-Bradley, 1948 (subsequent type designation)
Checked: verified by a taxonomic editorTrachyleberis Brady, 1898 (subsequent type designation)
Checked: verified by a taxonomic editorTrachyleberis (Acanthocythereis) (Howe, 1963) Siddiqui, 1971 † accepted as Checked: verified by a taxonomic editorAcanthocythereis Howe, 1963 (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Acanthocythereis) araneosa (Howe, 1963) Siddiqui, 1971 † accepted as Checked: verified by a taxonomic editorAcanthocythereis araneosa Howe, 1963 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Acanthocythereis) decoris Siddiqui, 1971 † accepted as Checked: verified by a taxonomic editorAcanthocythereis decoris (Siddiqui, 1971) Siddiqui, 1980 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Acanthocythereis) pedigaster Siddiqui, 1971 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Acanthocythereis) postcornis Siddiqui, 1971 † accepted as Checked: verified by a taxonomic editorAcanthocythereis postcornis (Siddiqui, 1971) Ahmad, Neale & Siddiqui, 1991 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Acanthocythereis) procapsus Siddiqui, 1971 † accepted as Checked: verified by a taxonomic editorAcanthocythereis procapsus (Siddiqui, 1971) Bhandari, 1992 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Acanthocythereis) usitata Siddiqui, 1971 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Actinocythereis) (Puri, 1953) Morkhoven, 1963 accepted as Checked: verified by a taxonomic editorActinocythereis Puri, 1953 (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Actinocythereis) birmanica Gramann, 1975 † accepted as Checked: verified by a taxonomic editorActinocythereis birmanica (Gramann, 1975) Hanai, Ikeya & Yajima, 1980 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Actinocythereis) birmanica birmanica Gramann, 1975 † accepted as Checked: verified by a taxonomic editorActinocythereis birmanica birmanica (Gramann, 1975) Hanai, Ikeya & Yajima, 1980 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Actinocythereis) birmanica kyaukkokensis Gramann, 1975 † accepted as Checked: verified by a taxonomic editorActinocythereis birmanica kyaukkokensis (Gramann, 1975) Hanai, Ikeya & Yajima, 1980 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Actinocythereis) birmanica pyawbwe Gramann, 1975 † accepted as Checked: verified by a taxonomic editorActinocythereis birmanica pyawbwe (Gramann, 1975) Hanai, Ikeya & Yajima, 1980 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Actinocythereis) exanthemata (Ulrich & Bassler, 1904) Morkhoven, 1963 † accepted as Checked: verified by a taxonomic editorActinocythereis exanthemata (Ulrich & Bassler, 1904) Puri, 1953 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Actinocythereis) marchilensis Hartmann, 1965 (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Actinocythereis) scutigera (Brady, 1868) Gramann, 1975 accepted as Checked: verified by a taxonomic editorActinocythereis scutigera (Brady, 1868) Goerlich, 1956 (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Actinocythereis) tumefacentis (Luebimova & Guha, 1960) Krstic, 1979 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Actinocythereis) wechesensis (Sutton & Williams, 1940) Morkhoven, 1963 † accepted as Checked: verified by a taxonomic editorActinocythereis wechesensis (Sutton & Williams, 1940) Puri, 1953 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Actinocythereis) woodwardsensis (Howe & Law, 1936) Morkhoven, 1963 † accepted as Checked: verified by a taxonomic editorActinocythereis woodwardsensis (Howe & Law, 1936) Puri, 1953 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Alteratrachyleberis) Li, 1964 accepted as Checked: verified by a taxonomic editorAlteratrachyleberis (Li, 1964) Liebau, 1977 † (new combination reference)
Checked: verified by a taxonomic editorTrachyleberis (Alteratrachyleberis) quasiperforata Kulichenko, 1976 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Costa) (Neviani, 1928) Keij, 1957 † accepted as Checked: verified by a taxonomic editorCosta (Neviani, 1928) Howe, 1955 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Costa) costata Scheremeta, 1969 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Costa) edwardsii (Roemer, 1838) Keij, 1957 † accepted as Checked: verified by a taxonomic editorCosta edwardsii (Roemer, 1838) † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Costa) elongata Scheremeta, 1969 † accepted as Checked: verified by a taxonomic editorMuellerina elongata (Scheremeta, 1969) Szczechura, 1977 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Echinocythereis) (Puri, 1954) Li, 1966 † accepted as Checked: verified by a taxonomic editorEchinocythereis Puri, 1954 (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Echinocythereis) scabra (Muenster, 1830) Li, 1966 accepted as Checked: verified by a taxonomic editorEchinocythereis scabra (Muenster, 1830) Oertli, 1956 (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Henryhowella) (Puri, 1957) Li, 1966 accepted as Checked: verified by a taxonomic editorHenryhowella Puri, 1957 (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Hermanites) (Puri, 1955) Li, 1966 accepted as Checked: verified by a taxonomic editorHermanites Puri, 1955 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Hermanites) bassleri (Ulrich, 1901) Li, 1966 † accepted as Checked: verified by a taxonomic editorHermanites bassleri (Ulrich, 1901) Fitz-gerald, 1983 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Hermanites) scribrosa (Li, 1963) Li, 1966 † accepted as Checked: verified by a taxonomic editorHermanites scribrosa (Li, 1963) † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Hermanites) suzakensis daedalia (Li, 1963) Li, 1966 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Hermanites) suzakensis suzakensis (Mandelstam, 1959) Li, 1966 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Hermanites) torosa (Li, 1963) Li, 1966 † accepted as Checked: verified by a taxonomic editorHermanites torosa (Li, 1963), 1966 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Rectotrachyleberis) Ruggieri, 1952 accepted as Checked: verified by a taxonomic editorRectotrachyleberis (Ruggieri, 1952) Sylvester-Bradley & Harding, 1954 (new combination reference)
Checked: verified by a taxonomic editorTrachyleberis (Rectotrachyleberis) edwardsii (Roemer, 1838) Ruggieri, 1952 † accepted as Checked: verified by a taxonomic editorCosta edwardsii (Roemer, 1838) † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Rectotrachyleberis) edwardsii edwardsii (Roemer, 1838) Ruggieri, 1952 † accepted as Checked: verified by a taxonomic editorCosta edwardsii edwardsii (Roemer, 1838) Keij, 1957 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Rectotrachyleberis) edwardsii padana (Ruggieri, 1950) Ruggieri, 1952 accepted as Checked: verified by a taxonomic editorCosta edwardsii padana (Ruggieri, 1950) † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Rectotrachyleberis) hamata (Mueller, 1894) Ruggieri, 1952 accepted as Checked: verified by a taxonomic editorCarinocythereis hamata (Mueller, 1894) Khosla, Mathur & Pant, 1982 (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Rectotrachyleberis) polytrema (Brady, 1878) Ruggieri, 1952 accepted as Checked: verified by a taxonomic editorCosta polytrema (Brady, 1878) Ruggieri, 1959 (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Rectotrachyleberis) tricostata (Reuss, 1850) Ruggieri, 1952 † accepted as Checked: verified by a taxonomic editorCosta (Cuneocosta) tricostata (Reuss, 1850) Ruggieri, 1992 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Trachyleberidea) (Bowen, 1953) Li, 1966 accepted as Checked: verified by a taxonomic editorTrachyleberidea Bowen, 1953 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Trachyleberis) Brady, 1898 (new combination reference)
Checked: verified by a taxonomic editorTrachyleberis (Trachyleberis) aculeata (Bosquet, 1852) Apostolescu, 1955 † accepted as Checked: verified by a taxonomic editorHorrificiella aculeata (Bosquet, 1852) Guernet, 1984 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Trachyleberis) arachnoidea (Bosquet, 1852) Keij, 1957 † represented as Checked: verified by a taxonomic editorTrachyleberis arachnoidea (Bosquet, 1852) Keij, 1957 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Trachyleberis) asperrima (Reuss, 1850) Keij, 1957 † accepted as Checked: verified by a taxonomic editorHenryhowella asperrima (Reuss, 1850) Bold, 1960 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Trachyleberis) asperrima asperrima (Reuss, 1850) Keij, 1957 † accepted as Checked: verified by a taxonomic editorHenryhowella asperrima asperrima (Reuss, 1850) Bold, 1960 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Trachyleberis) asperrima echinata (Reuss, 1851) Keij, 1957 † accepted as Checked: verified by a taxonomic editorTrachyleberis echinata (Reuss, 1851) Keij, 1957 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Trachyleberis) bimammillata Siddiqui, 1971 † represented as Checked: verified by a taxonomic editorTrachyleberis bimammillata Siddiqui, 1971 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Trachyleberis) dunelmensis (Norman, 1865) Morkhoven, 1963 accepted as Checked: verified by a taxonomic editorActinocythereis dunelmensis (Norman, 1865) Ruggieri, 1977 (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Trachyleberis) echinata (Reuss, 1851) Keij, 1957 † represented as Checked: verified by a taxonomic editorTrachyleberis echinata (Reuss, 1851) Keij, 1957 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Trachyleberis) hystrix (Reuss, 1850) Ruggieri, 1950 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Trachyleberis) lobuculus Siddiqui, 1971 † represented as Checked: verified by a taxonomic editorTrachyleberis lobuculus Siddiqui, 1971 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Trachyleberis) lyelliana (Bosquet, 1852) Keij, 1957 † accepted as Checked: verified by a taxonomic editorEchinocythereis lyelliana (Bosquet, 1852) Ellermann, 1960 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis (Trachyleberis) scabrocuneata (Brady, 1880) Brady, 1898 represented as Checked: verified by a taxonomic editorTrachyleberis scabrocuneata (Brady, 1880) Brady, 1898 (new combination reference)
Checked: verified by a taxonomic editorTrachyleberis (Trachyleberis) spinosa (Lienenklaus, 1900) Keij, 1957 † accepted as Checked: verified by a taxonomic editorAcanthocythereis spinosa (Lienenklaus, 1900) Moos, 1968 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis acanthoderma (Brady, 1880) Ruan & Hao (Yi-Chun), 1988 accepted as Checked: verified by a taxonomic editorLegitimocythere acanthoderma (Brady, 1880) Coles & Whatley, 1989 (additional source)
Checked: verified by a taxonomic editorTrachyleberis aculeata Rosyjeva, 1962 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis aculeata (Bosquet, 1852) Apostolescu, 1955 † accepted as Checked: verified by a taxonomic editorHorrificiella aculeata (Bosquet, 1852) Guernet, 1984 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis aculeata (Bosquet, 1852) Deroo, 1966 † accepted as Checked: verified by a taxonomic editorHorrificiella aculeata (Bosquet, 1852) Guernet, 1984 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis aculeata aculeata (Bosquet, 1852) Apostolescu, 1955 † accepted as Checked: verified by a taxonomic editorHorrificiella aculeata aculeata (Bosquet, 1852) Guernet, 1984 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis aculeata horridula (Bosquet, 1854) Margerie, 1968 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis aculeata minituberculata Margerie, 1968 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis aculeata modesta Van Hinte, 1962 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis acuminata Holden, 1964 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis adornata Chochlova, 1968 † accepted as Checked: verified by a taxonomic editorEchinocythereis adornata (Chochlova, 1968) Nikolaeva, 1971 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis anteplana Bate, 1972 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis arachnoidea (Bosquet, 1852) Keij, 1957 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis artschmanensis Rosyjeva, 1962 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis ascolii Puri, 1963 accepted as Checked: verified by a taxonomic editorAcanthocythereis ascolii (Puri, 1963) Bonaduce, Ciampo & Masoli, 1976 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis asperrima (Reuss, 1850) Keij, 1957 † accepted as Checked: verified by a taxonomic editorHenryhowella asperrima (Reuss, 1850) Bold, 1960 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis asperrima asperrima (Reuss, 1850) Keij, 1957 † accepted as Checked: verified by a taxonomic editorHenryhowella asperrima asperrima (Reuss, 1850) Bold, 1960 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis asperrima echinata (Reuss, 1851) Keij, 1957 † accepted as Checked: verified by a taxonomic editorTrachyleberis echinata (Reuss, 1851) Keij, 1957 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis avitis (Schneider, 1953) Uljanova, 1965 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis babadurmasensis Rosyjeva, 1962 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis bahamensis (Brady, 1870) Puri, 1953 (additional source)
Checked: verified by a taxonomic editorTrachyleberis barbata Gramm, 1963 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis barbatus Gramm, 1963 † accepted as Checked: verified by a taxonomic editorTrachyleberis barbata Gramm, 1963 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis baruni Banerji, 1970 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis bassleri (Ulrich, 1901) Brown, 1958 † accepted as Checked: verified by a taxonomic editorHermanites bassleri (Ulrich, 1901) Fitz-gerald, 1983 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis bassleri (Ulrich, 1901) Swain, 1952 † accepted as Checked: verified by a taxonomic editorHermanites bassleri (Ulrich, 1901) Fitz-gerald, 1983 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis bathymarina Ayress, 1993 accepted as Checked: verified by a taxonomic editorAyressoleberis bathymarina (Ayress, 1993) Brandão & Yasuhara, 2013 (additional source)
Checked: verified by a taxonomic editorTrachyleberis baturini Schneider, 1959 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis bermudezi (Bold, 1946) Bold, 1960 † accepted as Checked: verified by a taxonomic editorLegitimocythere bermudezi (Bold, 1946) Coles & Whatley, 1989 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis bermudezi bermudezi (Bold, 1946) Bold, 1960 † accepted as Checked: verified by a taxonomic editorLegitimocythere bermudezi bermudezi (Bold, 1946) Coles & Whatley, 1989 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis bermudezi crebripustulosa Bold, 1966 † accepted as Checked: verified by a taxonomic editorLegitimocythere bermudezi crebripustulosa (Bold, 1966) Coles & Whatley, 1989 (additional source)
Checked: verified by a taxonomic editorTrachyleberis bhujensis Tewari & Tandon, 1960 † accepted as Checked: verified by a taxonomic editorAcanthocythereis bhujensis (Tewari & Tandon, 1960) Guha, 1968 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis bicarinata (Swain, 1946) Swain, 1952 † accepted as Checked: verified by a taxonomic editorQuadracythere bicarinata (Swain, 1946) Neufville, 1973 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis bimammillata Siddiqui, 1971 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis bodjonegoroensis (Kingma, 1948) Guha, 1968 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis bollii Bold, 1960 † accepted as Checked: verified by a taxonomic editorBathypterocythereis bollii (Bold, 1960) Coles & Whatley, 1989 † (additional source)
Checked: verified by a taxonomic editorTrachyleberis bowerbankiana (Jones, 1857) Bowen, 1953 † accepted as Checked: verified by a taxonomic editorAgrenocythere bowerbankiana (Jones, 1857) King, 1981 † (additional source)

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2013-10-20 19:45:18Z  created  Brandão, Simone Nunes
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