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Pinkster S. (1993). A Revision of the genus Echinogammarus Stebbing, 1899 with some notes on related genera (Crustacea, Amphipoda). Memorie del Museo Civico di Storia Naturale, Verona (ser 2). 10, 1-185.
15680
Pinkster S.
1993
A Revision of the genus Echinogammarus Stebbing, 1899 with some notes on related genera (Crustacea, Amphipoda)
Memorie del Museo Civico di Storia Naturale, Verona (ser 2)
10, 1-185.
Publication
Available for editors  PDF available [request]
The taxonomic status of the genus Echinogammarus is discussed in relation to a great number of related genera. The type material or topotypic material of all studied taxa was (re)examined (if still existing) and compared with many (about 4000) newly collected samples and material from other museums and institutes in Europe. The morphological characters used in this work are identical to those used by Karaman & Pinkster, 1977 a, b; 1987. The genera Chaetogammarus, Marinogammarus, Ostiogammarus, Homoeogammarus, and Parhomoeogammarus are considered synonyms of Echinogammarus. Unlike Karaman, 1977 a, b, and Barnard & Barnard, 1983 the genera Pectenogammarus, Fontogammarus, Sarothrogammarus, Neogammarus, Rhipidogammarus, Longigammarus, and Eulimnogammarus are preserved as independent genera. This is mainly based on morphological differences in the mouthparts. The characters of Echinogammarus are discussed and a new definition of the genus is given. Complete descriptions and many illustrations of males (and if necessary of females) are given of all species known with certainty from Europe, Asia Minor and northern Africa. Five new species are described: E. kretensis, E. sardus, E. reductus, E. kerkuraios, and E. platvoeti. One subspecies is raised to the specific rank: E. monomerus. The variability of the various characters (both seasonal and geographical) is discussed and a key is given of both the Echinogammarus generacomplex and the (sub)species of the genus Echinogammarus. If available, information about the ecology of the various taxa is given.
Freshwater
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Echinogammarus acarinatus (S. Karaman, 1931) accepted as Dinarogammarus acarinatus (S. Karaman, 1931) (additional source)
Echinogammarus adipatus Karaman & Tibaldi, 1972 accepted as Echinogammarus incertae sedis adipatus Karaman & Tibaldi, 1972 (additional source)
Echinogammarus afer Stock, 1974 accepted as Echinogammarus incertae sedis afer Stock, 1974 (additional source)
Echinogammarus annandalei (Monod, 1924) accepted as Echinogammarus incertae sedis annandalei (Monod, 1924) (additional source)
Echinogammarus antalyae G. Karaman, 1971 accepted as Pectenogammarus antalyae (G. S. Karaman, 1971) (additional source)
Echinogammarus apfelbecki (S. Karaman, 1931) accepted as Echinogammarus incertae sedis apfelbecki (S. Karaman, 1931) (additional source)
Echinogammarus aquilifer Pinkster, 1969 (additional source)
Echinogammarus berilloni (Catta, 1878) (additional source)
Echinogammarus bolo Karaman & Tibaldi, 1972 accepted as Echinogammarus tibaldii Pinkster & Stock, 1970 accepted as Pectenogammarus tibaldii (Pinkster & Stock, 1970) (source of synonymy)
Echinogammarus cari (S. Karaman, 1931) accepted as Dinarogammarus cari (S. Karaman, 1931) (additional source)
Echinogammarus cyrtus Pinkster & Platvoet, 1986 accepted as Pectenogammarus cyrtus (Pinkster & Platvoet, 1986) (additional source)
Echinogammarus dactylus G. Karaman, 1987 accepted as Echinogammarus incertae sedis dactylus G. Karaman, 1987 (additional source)
Echinogammarus echinosetosus Pinkster, 1973 accepted as Echinogammarus incertae sedis echinosetosus Pinkster, 1973 (additional source)
Echinogammarus eisentrauti (Schellenberg, 1937) accepted as Echinogammarus incertae sedis eisentrauti (Schellenberg, 1937) (additional source)
Echinogammarus feminatus Pinkster, 1973 (additional source)
Echinogammarus fluminensis Pinkster & Stock, 1970 accepted as Echinogammarus stammeri (S. Karaman, 1931) accepted as Pectenogammarus stammeri (S. Karaman, 1931) (additional source)
Echinogammarus incertae sedis tacapensis (Chevreux & Gauthier, 1924) (taxonomy source)
Echinogammarus kerkuraios Pinkster, 1993 accepted as Echinogammarus incertae sedis kerkuraios Pinkster, 1993 (original description)
Echinogammarus klaptoczi Schäferna, 1908 accepted as Echinogammarus incertae sedis klaptoczi Schäferna, 1908 (additional source)
Echinogammarus kretensis Pinkster, 1993 accepted as Echinogammarus incertae sedis klaptoczi Schäferna, 1908 (original description)
Echinogammarus macrocarpus (Stock, 1969) accepted as Iberogammarus macrocarpus (Stock, 1969) (source of synonymy)
Echinogammarus margalefi Pinkster, 1973 accepted as Echinogammarus incertae sedis margalefi Pinkster, 1973 (additional source)
Echinogammarus meridionalis Pinkster, 1973 accepted as Echinogammarus incertae sedis meridionalis Pinkster, 1973 (additional source)
Echinogammarus monomerus Stock, 1978 accepted as Pectenogammarus monomerus (Stock, 1978) (redescription)
Echinogammarus pacaudi Hubault & Ruffo, 1956 accepted as Parhomoeogammarus pacaudi (Hubault & Ruffo, 1956) (additional source)
Echinogammarus paxjuliensis Mateus, Mateus & Afonso, 1979 accepted as Echinogammarus simoni (Chevreux, 1894) accepted as Pectenogammarus simoni (Chevreux, 1894) (additional source)
Echinogammarus pinksteri Van Maren, 1973 accepted as Echinogammarus incertae sedis pinksteri Van Maren, 1973 (additional source)
Echinogammarus pirloti (Sexton & Spooner, 1940) accepted as Marinogammarus pirloti Sexton & Spooner, 1940 (additional source)
Echinogammarus platvoeti Pinkster, 1993 accepted as Pectenogammarus platvoeti (Pinkster, 1993) (original description)
Echinogammarus pungens (H. Milne Edwards, 1840) accepted as Pectenogammarus pungens (H. Milne Edwards, 1840) (additional source)
Echinogammarus pungens carinata (Pljakić, 1962) accepted as Echinogammarus pungens (H. Milne Edwards, 1840) accepted as Pectenogammarus pungens (H. Milne Edwards, 1840) (additional source)
Echinogammarus pungens minoricensis (Margalef, 1952) accepted as Echinogammarus pungens (H. Milne Edwards, 1840) accepted as Pectenogammarus pungens (H. Milne Edwards, 1840) (additional source)
Echinogammarus pungentoides Diviacco & Pinkster, 1982 accepted as Pectenogammarus pungens (H. Milne Edwards, 1840) (source of synonymy)
Echinogammarus rashiini Alouf, 1976 accepted as Echinogammarus incertae sedis rashiini Alouf, 1976 (additional source)
Echinogammarus reductus Pinkster, 1993 accepted as Echinogammarus incertae sedis reductus Pinkster, 1993 (original description)
Echinogammarus roco G. Karaman, 1973 accepted as Echinogammarus tibaldii Pinkster & Stock, 1970 accepted as Pectenogammarus tibaldii (Pinkster & Stock, 1970) (additional source)
Echinogammarus ruffoi Pinkster & Stock, 1970 accepted as Pectenogammarus ruffoi (Pinkster & Stock, 1970) (additional source)
Echinogammarus sardus Pinkster, 1993 accepted as Echinogammarus incertae sedis sardus Pinkster, 1993 (original description)
Echinogammarus sicilianus Karaman & Tibaldi, 1972 accepted as Echinogammarus incertae sedis sicilianus Karaman & Tibaldi, 1972 (additional source)
Echinogammarus simoni (Chevreux, 1894) accepted as Pectenogammarus simoni (Chevreux, 1894) (additional source)
Echinogammarus spinulicornis Pinkster & Stock, 1971 accepted as Pectenogammarus spinulicornis (Pinkster & Stock, 1971) (additional source)
Echinogammarus stammeri (S. Karaman, 1931) accepted as Pectenogammarus stammeri (S. Karaman, 1931) (additional source)
Echinogammarus stammeri stammeri G. Karaman, 1974 accepted as Echinogammarus stammeri (S. Karaman, 1931) accepted as Pectenogammarus stammeri (S. Karaman, 1931) (additional source)
Echinogammarus stammeri visualis S. Karaman, 1974 accepted as Echinogammarus stammeri (S. Karaman, 1931) accepted as Pectenogammarus stammeri (S. Karaman, 1931) (additional source)
Echinogammarus stocki G. Karaman, 1970 accepted as Pectenogammarus stocki (G. Karaman, 1970) (additional source)
Echinogammarus tabu G. Karaman, 1971 accepted as Pectenogammarus tabu (G. Karaman, 1971) (additional source)
Echinogammarus tabu mutus Karaman & Tibaldi, 1972 (additional source)
Echinogammarus tabu tabu G. Karaman, 1971 represented as Echinogammarus tabu G. Karaman, 1971 accepted as Pectenogammarus tabu (G. Karaman, 1971) (additional source)
Echinogammarus tacapensis (Chevreux & Gauthier, 1924) accepted as Echinogammarus incertae sedis tacapensis (Chevreux & Gauthier, 1924) (additional source)
Echinogammarus tarragonensis Pinkster, 1973 (additional source)
Echinogammarus thoni (Schäferna, 1923) accepted as Pectenogammarus thoni (Schäferna, 1923) (additional source)
Echinogammarus tibaldii Pinkster & Stock, 1970 accepted as Pectenogammarus tibaldii (Pinkster & Stock, 1970) (additional source)
Echinogammarus valedictus Platvoet & Pinkster, 1990 accepted as Echinogammarus incertae sedis valedictus Platvoet & Pinkster, 1990 (additional source)
Echinogammarus veneris (Heller, 1865) accepted as Pectenogammarus veneris (Heller, 1865) (additional source)
Echinogammarus zebrinus Pinkster & Stock, 1971 (additional source)
Gammarus finmarchicus Dahl, 1938 accepted as Echinogammarus finmarchicus (Dahl, 1938) accepted as Echinogammarus incertae sedis finmarchicus (Dahl, 1938) (source of synonymy)
Gammarus lochites Margalef, 1956 accepted as Echinogammarus lochites (Margalef, 1956) accepted as Echinogammarus incertae sedis lochites (Margalef, 1956) (additional source)
Gammarus pungens H. Milne Edwards, 1840 accepted as Echinogammarus pungens (H. Milne Edwards, 1840) accepted as Pectenogammarus pungens (H. Milne Edwards, 1840) (additional source)
Gammarus simoni Chevreux, 1894 accepted as Echinogammarus simoni (Chevreux, 1894) accepted as Pectenogammarus simoni (Chevreux, 1894) (additional source)
Gammarus tacapensis Chevreux & Gauthier, 1924 accepted as Echinogammarus tacapensis (Chevreux & Gauthier, 1924) accepted as Echinogammarus incertae sedis tacapensis (Chevreux & Gauthier, 1924) (additional source)
Laurogammarus scutarensis (Schäferna, 1923) accepted as Pectenogammarus scutarensis (Schäferna, 1923) (identification resource)
Ostiogammarus acarinatus stammeri S. Karaman, 1931 accepted as Echinogammarus stammeri (S. Karaman, 1931) accepted as Pectenogammarus stammeri (S. Karaman, 1931) (additional source)
Ostiogammarus pungens (H. Milne Edwards, 1840) accepted as Echinogammarus pungens (H. Milne Edwards, 1840) accepted as Pectenogammarus pungens (H. Milne Edwards, 1840) (additional source)
Algeria for Echinogammarus annandalei (Monod, 1924) 
Algeria for Echinogammarus reductus Pinkster, 1993 
Algeria for Echinogammarus simoni (Chevreux, 1894) 
Algeria for Echinogammarus tacapensis (Chevreux & Gauthier, 1924) 
Algeria for Echinogammarus valedictus Platvoet & Pinkster, 1990 
Algeria for Echinogammarus valedictus Platvoet & Pinkster, 1990 
Baleares for Echinogammarus eisentrauti (Schellenberg, 1937) 
Baleares for Echinogammarus klaptoczi Schäferna, 1908 
Baleares for Echinogammarus klaptoczi Schäferna, 1908 
Baleares for Echinogammarus pungens (H. Milne Edwards, 1840) 
Belgian Exclusive Economic Zone for Echinogammarus berilloni (Catta, 1878) 
Bosnia and Herzegovina for Echinogammarus acarinatus (S. Karaman, 1931) 
Corse for Echinogammarus stocki G. Karaman, 1970 
Croatia for Echinogammarus acarinatus (S. Karaman, 1931) 
Croatia for Echinogammarus cari (S. Karaman, 1931) 
Cyprus for Echinogammarus veneris (Heller, 1865) 
Dutch Exclusive Economic Zone for Echinogammarus berilloni (Catta, 1878) 
France for Echinogammarus acarinatus (S. Karaman, 1931)  (origin: native - endemic)
France for Echinogammarus aquilifer Pinkster, 1969 
France for Echinogammarus berilloni (Catta, 1878) 
France for Echinogammarus cyrtus Pinkster & Platvoet, 1986 
France for Echinogammarus feminatus Pinkster, 1973 
France for Echinogammarus pacaudi Hubault & Ruffo, 1956 
France for Echinogammarus pungens (H. Milne Edwards, 1840) 
France for Echinogammarus spinulicornis Pinkster & Stock, 1971 
France for Echinogammarus stammeri (S. Karaman, 1931) 
France for Echinogammarus stocki G. Karaman, 1970 
France for Echinogammarus tabu G. Karaman, 1971 
France for Echinogammarus tabu tabu G. Karaman, 1971 
Germany for Echinogammarus berilloni (Catta, 1878) 
Greece for Echinogammarus apfelbecki (S. Karaman, 1931) 
Greece for Echinogammarus kerkuraios Pinkster, 1993 
Greece for Echinogammarus pinksteri Van Maren, 1973 
Greece for Echinogammarus platvoeti Pinkster, 1993 
Greece for Echinogammarus platvoeti Pinkster, 1993 
Greece for Echinogammarus pungens (H. Milne Edwards, 1840) 
Greece for Echinogammarus veneris (Heller, 1865) 
Italy for Echinogammarus pinksteri Van Maren, 1973 
Italy for Echinogammarus pungens (H. Milne Edwards, 1840) 
Italy for Echinogammarus stammeri (S. Karaman, 1931) 
Italy for Echinogammarus stocki G. Karaman, 1970 
Italy for Echinogammarus tibaldii Pinkster & Stock, 1970 
Italy for Echinogammarus tibaldii Pinkster & Stock, 1970 
Italy for Echinogammarus tibaldii Pinkster & Stock, 1970 
Italy for Echinogammarus tibaldii Pinkster & Stock, 1970 
Italy for Echinogammarus tibaldii Pinkster & Stock, 1970 
Italy for Echinogammarus tibaldii Pinkster & Stock, 1970 
Italy for Echinogammarus veneris (Heller, 1865) 
Kriti for Echinogammarus kretensis Pinkster, 1993 
Kriti for Echinogammarus veneris (Heller, 1865) 
Lebanon-Syria for Echinogammarus veneris (Heller, 1865) 
Libya for Echinogammarus afer Stock, 1974 
Libya for Echinogammarus klaptoczi Schäferna, 1908 
Libya for Echinogammarus klaptoczi Schäferna, 1908 
Montenegro for Echinogammarus acarinatus (S. Karaman, 1931) 
Morocco for Echinogammarus simoni (Chevreux, 1894) 
Palestine for Echinogammarus veneris (Heller, 1865) 
Portugal for Echinogammarus meridionalis Pinkster, 1973 
Portugal for Echinogammarus simoni (Chevreux, 1894) 
Sardegna for Echinogammarus pungens (H. Milne Edwards, 1840) 
Sardegna for Echinogammarus sardus Pinkster, 1993 
Sardegna for Echinogammarus sicilianus Karaman & Tibaldi, 1972 
Sardegna for Echinogammarus tibaldii Pinkster & Stock, 1970 
Sardegna for Echinogammarus tibaldii Pinkster & Stock, 1970 
Sardegna for Echinogammarus tibaldii Pinkster & Stock, 1970 
Sardegna for Echinogammarus veneris (Heller, 1865) 
Sardinia for Echinogammarus adipatus Karaman & Tibaldi, 1972 
Sicilia for Echinogammarus sicilianus Karaman & Tibaldi, 1972 
Sicilia for Echinogammarus sicilianus Karaman & Tibaldi, 1972 
Sicilia for Echinogammarus sicilianus Karaman & Tibaldi, 1972 
Sicilia for Echinogammarus tibaldii Pinkster & Stock, 1970 
Sicilia for Echinogammarus tibaldii Pinkster & Stock, 1970 
Sicilia for Echinogammarus tibaldii Pinkster & Stock, 1970 
Sicilia for Echinogammarus veneris (Heller, 1865) 
Sicily for Echinogammarus adipatus Karaman & Tibaldi, 1972 
Sicily for Echinogammarus klaptoczi Schäferna, 1908 
Spain for Echinogammarus aquilifer Pinkster, 1969 
Spain for Echinogammarus berilloni (Catta, 1878) 
Spain for Echinogammarus echinosetosus Pinkster, 1973 
Spain for Echinogammarus feminatus Pinkster, 1973 
Spain for Echinogammarus lochites (Margalef, 1956) 
Spain for Echinogammarus longisetosus Pinkster, 1973 
Spain for Echinogammarus margalefi Pinkster, 1973 
Spain for Echinogammarus monomerus Stock, 1978 
Spain for Echinogammarus navaensis Lop, 1987 
Spain for Echinogammarus obtusidens Pinkster & Stock, 1972 
Spain for Echinogammarus obtusidens Pinkster & Stock, 1972 
Spain for Echinogammarus pacaudi Hubault & Ruffo, 1956 
Spain for Echinogammarus pseudoaquilifer Platvoet & Pinkster, 1980 
Spain for Echinogammarus pungens (H. Milne Edwards, 1840) 
Spain for Echinogammarus simoni (Chevreux, 1894) 
Switzerland for Echinogammarus berilloni (Catta, 1878) 
Switzerland for Echinogammarus stammeri (S. Karaman, 1931) 
Tunisia for Echinogammarus afer Stock, 1974 
Tunisia for Echinogammarus dactylus G. Karaman, 1987 
Tunisia for Echinogammarus simoni (Chevreux, 1894) 
Tunisia for Echinogammarus tacapensis (Chevreux & Gauthier, 1924) 
Turkey for Echinogammarus antalyae G. Karaman, 1971 
Turkey for Echinogammarus stocki G. Karaman, 1970 
Turkey for Echinogammarus veneris (Heller, 1865)