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Register of
Marine Species
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Pennella Oken, 1815

135648 (urn:lsid:marinespecies.org:taxname:135648)

  1. Biota
  2. Animalia (Kingdom)
  3. Arthropoda (Phylum)
  4. Crustacea (Subphylum)
  5. Multicrustacea (Superclass)
  6. Copepoda (Class)
  7. Neocopepoda (Infraclass)
  8. Podoplea (Superorder)
  9. Siphonostomatoida (Order)
  10. Pennellidae (Family)
  11. Pennella (Genus)
accepted
Genus
Pennellidae Burmeister, 1835
marine, brackish, fresh, terrestrial
Oken, L. (1815 - 1816). Lehrbuch der Naturgeschichte. Dritter Theil: Zoologie. [Textbook of Natural History. Part Three: Zoology.]. <em>Erste Abtheilung: Fleischlose Thiere,Leipzig: C.H. Reclam & Jena: A. Schmid, [book (3rd vol part one, of 3 vols, including plates atlas of T.1, 1813].</em> xxviii + 842 pp. + xviii, 40 pls. L [copepods 180-184, 357-359, 4 plates]., available online at https://doi.org/10.5962/bhl.title.166403 [details] OpenAccess publication
Walter, T.C.; Boxshall, G. (2021). World of Copepods database. Pennella Oken, 1815. Accessed through: Xu, K., An, J., Ding, L., Dong, D., Gao, Y., Huang, Y., Jiang, W., Lei, Y., Li, K., Li, X., Li, Y., Liang, X., Lin, G., Lin, M., Liu, H., Liu, J., Liu, W., Liu, X., Lu, D., Ma, Z., Ren, X., Sha, Z., Sun, J., Sun, S., Sun, Z., Tian, L., Wang, D., Wang, J., Wu, X., Xia, B., Xiao, N., Yin, J., Zhang, J., Zhang, S., Zhang, X., Zhao, E., Zheng, X., Zhou, H., Zhou, J. & Zhou, K. (2021) Chinese Register of Marine Species at: https://www.marinespecies.org/charms/aphia.php?p=taxdetails&id=135648 on 2026-08-17
Xu, K., An, J., Ding, L., Dong, D., Gao, Y., Huang, Y., Jiang, W., Lei, Y., Li, K., Li, X., Li, Y., Liang, X., Lin, G., Lin, M., Liu, H., Liu, J., Liu, W., Liu, X., Lu, D., Ma, Z., Ren, X., Sha, Z., Sun, J., Sun, S., Sun, Z., Tian, L., Wang, D., Wang, J., Wu, X., Xia, B., Xiao, N., Yin, J., Zhang, J., Zhang, S., Zhang, X., Zhao, E., Zheng, X., Zhou, H., Zhou, J. & Zhou, K. (2026). Chinese Register of Marine Species. Pennella Oken, 1815. Accessed at: https://www.marinespecies.org/charms/aphia.php?p=taxdetails&id=135648 on 2026-08-17
Date
action
by
2004-12-21 15:54:05Z
created
Boxshall, Geoff
2006-09-27 07:06:07Z
changed
Martinez, Olga
2008-08-06 13:32:15Z
changed
Walter, T. Chad
2011-04-05 08:08:50Z
changed
Boxshall, Geoff
2020-10-26 12:27:51Z
changed
Walter, T. Chad

original description Oken, L. (1815 - 1816). Lehrbuch der Naturgeschichte. Dritter Theil: Zoologie. [Textbook of Natural History. Part Three: Zoology.]. <em>Erste Abtheilung: Fleischlose Thiere,Leipzig: C.H. Reclam & Jena: A. Schmid, [book (3rd vol part one, of 3 vols, including plates atlas of T.1, 1813].</em> xxviii + 842 pp. + xviii, 40 pls. L [copepods 180-184, 357-359, 4 plates]., available online at https://doi.org/10.5962/bhl.title.166403 [details] OpenAccess publication

basis of record Boxshall, G. (2001). Copepoda (excl. Harpacticoida), <B><I>in</I></B>: Costello, M.J. <i>et al.</i> (Ed.) (2001). <i>European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels,</i> 50: pp. 252-268 (look up in IMIS) [details] 

additional source Neave, S.A. (1939 - 1996). Nomenclator Zoologicus. vol. 1-10 Online., available online at https://www.checklistbank.org/dataset/126539/about [details] 

additional source Wilson, C.B. (1917). North American parasitic copepods belonging to the Lernaeidae with a revision of the entire family. <em>Proceedings of the United States National Museum.</em> 53(2194):1-150, pls. 1-21. (13-vi-1917)., available online at https://doi.org/10.5479/si.00963801.53-2194.1
page(s): 106-112 [details] OpenAccess publication

additional source International Commission on Zoological Nomenclature, Opinion. (1995). Clavella Oken, 1815 and Pennella Oken, 1815 (Crustacea, Copepoda): conserved, and Pennella diodontis Oken, 1815: Specific name conserved. <em>Bulletin of Zoological Nomenclature.</em> 52(3):279-280. [details] Available for editors  PDF available

additional source Gentry, A. (1994). Comments on the proposed conservation of Clavella Oken, 1815 and Pennella Oken, 1815 (Crustacea, Copepoda). Bulletin of Zoological Nomenclature 51(4):339., available online at https://doi.org/10.5962/bhl.part.7240 [details] 

additional source Fage, L. (1931). Remarques sur le parasitisme des Copépodes du genre Penella. [Remarks on the parasitism of copepods of the genus Penella.]. <em>Bulletin de la Société Zoologique de France.</em> 56:190-193. [details] Available for editors  PDF available

additional source Castro Romero, R. & H. Baeza Kuroki. (1986). Pre-metamorphosis stages of two pennellids (Copepoda, Siphonostomatoida) from their definite host. <em>Crustaceana (Leiden).</em> 50(2):166-175, figs. 1-23. (iii.1986)., available online at https://doi.org/10.1163/156854086x00188 [details] Available for editors  PDF available

additional source Kabata, Z. (1979). Parasitic Copepoda of British Fishes. <em>The Ray Society, London.</em> 152: i-xii, 1-468, figs. 1-203. [details] Available for editors  PDF available

additional source Kabata, Z. (1988). Copepoda and Brachiura. In: Margolis, L. & Z. Kabata (eds.). Guide to the parasites of fishes of Canada. Part II. Crustacea. Canadian Special Publication in Fisheries and Aquatic Sciences 101:3-127, figs. 1-53. (French summary). [details] Available for editors  PDF available

additional source Kabata, Z. (1993). Clavella Oken, 1815 and Pennella Oken, 1815 (Crustacea, Copepoda): proposed conservation. Case 836. Bulletin of Zoological Nomenclature 50(4):273-276., available online at https://doi.org/10.5962/bhl.part.1872 [details] Available for editors  PDF available

additional source Lutken, C.F. (1893). Slaegten Baculus Lubbock, et Udviklingstrin af Pennella. [The genus Baculus Lubbock, a developmental stage of Pennella.]. <em>Videnskabelige Meddelelser fra Dansk Naturhistoriske Forening, Copenhagen.</em> (5)4:73-76. [details] 

additional source Rice, D.W. (1994). Comments on the proposed conservation of Clavella Oken, 1815 and Pennella Oken, 1815 (Crustacea, Copepoda). Bulletin of Zoological Nomenclature 51(4):338-339., available online at https://doi.org/10.5962/bhl.part.7240 [details] Available for editors  PDF available

additional source Pillai, N.K. (1985). Copepod Parasites of Marine Fishes. <em>In: The Fauna of India. Zoological Society of India, Calcutta.</em> 900pp., figs. 1-279.
page(s): 740 [details] Available for editors  PDF available

additional source Quidor, A. (1913). Copépodes Parasites. <em>In: Deuxième Expédition Antarctique Français (1908-1910). Sciences Naturelles, Documents Scientifiques.</em> (1908-1910): 197-214, pls. 1-4. [details] OpenAccess publication

additional source Rose, M. (1941). Sur le mâle de Pennella. Copépode marin parasite des Poissons et des Cétacés. [On the male of Pennella. Marine copepod parasitic on fish and cetaceans. <em>Comptes Rendus des Séances de la Société de Biologie, Paris.</em> 135(17-18):1478-1479. [details] Available for editors  PDF available

additional source Rose, M. (1943). II. Description d'un mâle du genre Pennella Oken, 1815. [II. Description of a male of the genus Pennella Oken, 1815.]. <em>Archives de Zoologie Expérimentale et Générale.</em> 83(1):22-33. [details] Available for editors  PDF available

additional source Hogans, W.E. (2017). Review of Pennella Oken, 1816 ( Copepoda: Pennellidae) with a description of Pennella benzi sp nov., a parasite of Escolar, Lepidocybium flavobrunneum (Pisces) in the northwest Atlantic Ocean. Zootaxa, 4244(1):1-38., available online at https://doi.org/10.11646/zootaxa.4244.1.1 [details] Available for editors  PDF available

additional source Suyama, S., Y. Masuda, T. Yanagimoto & S. Chow. (2019). Genetic and morphological variation in Pennella sp. (Copepoda: Siphonostomatoida) collected from Pacific saury, Cololabis saira. <em>Marine Biodiversity.</em> 49(3):1233-1245., available online at https://doi.org/10.1007/s12526-018-0901-x [details] Available for editors  PDF available

additional source Heldt, J.H. (1952). Note sur un male de Pennella (Copépoda parasite). <em>Bulletin de la Société des Sciences Naturelles de Tunisie.</em> 5:169-170. [details] Available for editors  PDF available

additional source Suyama, S., T. Yanagimoto, K. Nakai, T. Tamura, K. Shiozaki, S. Ohsimo & S. Chow. (2021). A taxonomic revision of Pennella Oken, 1815 based on morphology and genetics (Copepods: Siphonostomatoida: Pennellidae). <em>Journal of Crustacean Biology.</em> 41(3):1-12., available online at https://doi.org/10.1093/jcbiol/ruab040 [details] Available for editors  PDF available

additional source Mugetti, D., E. Colombino, V. Menconi, F. Garibaldi, W. Mignone, A. Gustinelli, M. Prearo, F. Guarda & M.T. Capucchio. (2021). Unusual localization of Pennella sp. in swordfish (Xiphias gladius) hearts. <em>Animals.</em> 11(6):1-5. Jun 2021., available online at https://doi.org/10.3390/ani11061757 [details] Available for editors  PDF available

additional source Ten, S., R.V. Pool, J.A. Raga, A.D. Sweet & F.J. Aznar. (2025). Phylogeography of Pennella (Copepoda: Siphonostomatoida: Pennellidae) indicates interoceanic dispersal mediated by cetacean and fish hosts. <em>Parasitology.</em> 152:1-10., available online at https://doi.org/10.1017/s0031182025000101 [details] Available for editors  PDF available
 
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3x5 C.B. Wilson Taxonomic Card - Pennella Document hosted within ChaRMS 
3x5 J.S. Ho Taxonomic Card - Pennella Document hosted within ChaRMS 
To Biological Information System for Marine Life (BISMaL) 
To European Nucleotide Archive, ENA (Pennella) 
To European Nucleotide Archive, ENA (Pennella) 
To Genbank 
To Genbank 
To Yale Peabody Museum of Natural History (YPM IZ 030291) 
To ITIS
Unreviewed
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