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Mantelatto, F.L.; Robles, R.; Schubart, C.D.; Felder, D.L. (2009). Molecular phylogeny of the genus Cronius Stimpson, 1860, with reassignment of C. tumidulus and several American species of Portunus to the genus Achelous De Haan, 1833 (Brachyura: Portunidae). In: Crustacean Issues 18: Decapod Crustacean Phylogenetics, Martin, J.W., Crandall, K.A. & Felder, D.L. (eds). CRC Press, England, pp. 567–579.