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Bouillon, J.; Boero, F. & Seghers, G. (1991). Notes additionelles sur les méduses de Papouasie Nouvelle-Guinée (Hydrozoa, Cnidaria) IV. Additional notes on the medusae of Papua New Guinea (Hydrozoa, Cnidaria): IV. Cah. Biol. mar. 32(3): 387-411.