WoRMS source details
Ekimova, I.; Carmona, L.; Mikhlina, A. L.; Grishina, D.; Stanovova, M. V.; Schepetov, D. M.; Hoover, C.; De Souza-Canal, J.; Kuznetsov, K. O.; Valdés, Á. (2026). Neither “lumpers” nor “splitters”: A global revision of Flabellinidae s.l. nudibranchs (Gastropoda: Heterobranchia: Nudibranchia). PLOS One. 21(5): e0347759.
563048
10.1371/journal.pone.0347759 [view]
Ekimova, I.; Carmona, L.; Mikhlina, A. L.; Grishina, D.; Stanovova, M. V.; Schepetov, D. M.; Hoover, C.; De Souza-Canal, J.; Kuznetsov, K. O.; Valdés, Á.
2026
Neither “lumpers” nor “splitters”: A global revision of Flabellinidae s.l. nudibranchs (Gastropoda: Heterobranchia: Nudibranchia)
PLOS One
21(5): e0347759
Publication
Stated date: 20 May 2026
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Apata Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Apata pricei (MacFarland, 1966) (status source)
Apataidae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Baenopsis Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Baenopsis baetica (García-Gómez, 1984) (status source)
Bajaeolis Gosliner & Behrens, 1986 (status source)
Bajaeolis bertschi Gosliner & Behrens, 1986 (status source)
Borealea Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella J. E. Gray, 1850 (source of synonymy)
Carronella Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Flabellina McMurtrie, 1831 (source of synonymy)
Chlamylla Bergh, 1886 (status source)
Chlamylla borealis Bergh, 1886 (status source)
Chlamylla ignicrystalla (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (new combination reference)
Chlamylla intermedia (Bergh, 1899) (status source)
Chlamylla islandica (Odhner, 1937) (new combination reference)
Chlamylla parva (M. Hadfield, 1963) (new combination reference)
Chlamylla polaris (Volodchenko, 1946) (new combination reference)
Corrupta Korshunova, Fletcher & Martynov, 2025 accepted as Coryphella J. E. Gray, 1850 (source of synonymy)
Coryphella J. E. Gray, 1850 (status source)
Coryphella abei Baba, 1987 (status source)
Coryphella alexanderi Ekimova, 2022 (status source)
Coryphella amabilis (Y. Hirano & Kuzirian, 1991) (status source)
Coryphella athadona Bergh, 1875 (status source)
Coryphella borealis Odhner, 1922 (status source)
Coryphella browni Picton, 1980 (status source)
Coryphella californica Bergh, 1904 (status source)
Coryphella candela (Korshunova, Fletcher & Martynov, 2025) (new combination reference)
Coryphella capensis Thiele, 1925 (status source)
Coryphella cerverai (M. A. Fischer, van der Velde & Roubos, 2007) (new combination reference)
Coryphella chriskaugei (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (status source)
Coryphella cooperi T. D. A. Cockerell, 1901 (status source)
Coryphella dana (Millen & Hamann, 2006) (new combination reference)
Coryphella falklandica Eliot, 1907 (status source)
Coryphella fogata (Millen & Hermosillo, 2007) (status source)
Coryphella gracilis (Alder & Hancock, 1844) (status source)
Coryphella insolita (García-Gómez & Cervera, 1989) (status source)
Coryphella lineata (Lovén, 1846) (status source)
Coryphella monicae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (status source)
Coryphella nobilis A. E. Verrill, 1880 (status source)
Coryphella orjani (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (status source)
Coryphella pallida A. E. Verrill, 1900 (status source)
Coryphella sanamyanae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (status source)
Coryphella trilineata O'Donoghue, 1921 (status source)
Coryphella trophina (Bergh, 1890) (status source)
Coryphellidae Bergh, 1889 accepted as Flabellinidae Bergh, 1889 (source of synonymy)
Coryphellina O'Donoghue, 1929 (status source)
Coryphellina albomarginata (M. C. Miller, 1971) (status source)
Coryphellina arveloi (Ortea & Espinosa, 1998) (additional source)
Coryphellina aurora Ekimova, Deart, Antokhina, Mikhlina & Schepetov, 2022 (status source)
Coryphellina delicata (Gosliner & Willan, 1991) (status source)
Coryphellina exoptata (Gosliner & Willan, 1991) (status source)
Coryphellina flamma Ekimova, Deart, Antokhina, Mikhlina & Schepetov, 2022 (status source)
Coryphellina hamanni (Gosliner, 1994) (status source)
Coryphellina indica (Bergh, 1902) (status source)
Coryphellina lotos Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Coryphellina marcusorum (Gosliner & Kuzirian, 1990) (status source)
Coryphellina pannae Ekimova, Deart, Antokhina, Mikhlina & Schepetov, 2022 (status source)
Coryphellina poenicia (Burn, 1957) (status source)
Coryphellina pseudolotos Ekimova, Deart, Antokhina, Mikhlina & Schepetov, 2022 (status source)
Coryphellina westralis (Burn, 1964) (status source)
Edmundsella Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Edmundsella albomaculata (Pola, Carmona, Calado & Cervera, 2014) (status source)
Edmundsella bertschi (Gosliner & Kuzirian, 1990) (status source)
Edmundsella dushia (Ev. Marcus & Er. Marcus, 1963) (new combination reference)
Edmundsella pedata (Montagu, 1816) (status source)
Edmundsella vansyoci (Gosliner, 1994) (status source)
Eolis rufibranchialis G. Johnston, 1832 accepted as Coryphella verrucosa (M. Sars, 1829) (source of synonymy)
Fjordia Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella J. E. Gray, 1850 (source of synonymy)
Flabellina bandeli (Ev. Marcus, 1976) (basis of record)
Flabellina cavolini (Vérany, 1846) (additional source)
Flabellina enne (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (basis of record)
Flabellina funeka Gosliner & R. J. Griffiths, 1981 (status source)
Flabellina gabinierei (Vicente, 1975) (basis of record)
Flabellina gaditana (Cervera, García-Gómez & F. J. García, 1987) (additional source)
Flabellina ischitana Y. Hirano & T. E. Thompson, 1990 (status source)
Flabellina pellucida (Alder & Hancock, 1843) (status source)
Flabellinidae Bergh, 1889 (status source)
Flabellinopsidae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Flabellinopsis MacFarland, 1966 (status source)
Flabellinopsis iodinea (J. G. Cooper, 1863) (status source)
Gulenia Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella J. E. Gray, 1850 (source of synonymy)
Kynaria Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Kynaria cynara (Ev. Marcus & Er. Marcus, 1967) (status source)
Launsina Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026 (original description)
Launsina rubropurpurata (Gosliner & Willan, 1991) (new combination reference)
Launsina tanyae Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026 (original description)
Luisella Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Luisella babai (Schmekel, 1972) (status source)
Luisella bulbosa (Ortea & Espinosa, 1998) (new combination reference)
Luisella engeli (Ev. Marcus & Er. Marcus, 1968) (new combination reference)
Luisella llerae (Ortea, 1989) (new combination reference)
Luisella telja (Ev. Marcus & Er. Marcus, 1967) (new combination reference)
Mgueolia Korshunova, Fletcher & Martynov, 2025 (status source)
Mgueolia anastasiae Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026 (original description)
Mgueolia ninae Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026 (original description)
Mgueolia vladimiri Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026 (original description)
Microchlamylla Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella J. E. Gray, 1850 (source of synonymy)
Notaeolidia Eliot, 1905 (status source)
Notaeolidia depressa Eliot, 1905 (status source)
Notaeolidia gigas Eliot, 1905 (status source)
Notaeolidia schmekelae Wägele, 1990 (status source)
Apata pricei (MacFarland, 1966) (status source)
Apataidae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Baenopsis Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Baenopsis baetica (García-Gómez, 1984) (status source)
Bajaeolis Gosliner & Behrens, 1986 (status source)
Bajaeolis bertschi Gosliner & Behrens, 1986 (status source)
Borealea Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella J. E. Gray, 1850 (source of synonymy)
Carronella Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Flabellina McMurtrie, 1831 (source of synonymy)
Chlamylla Bergh, 1886 (status source)
Chlamylla borealis Bergh, 1886 (status source)
Chlamylla ignicrystalla (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (new combination reference)
Chlamylla intermedia (Bergh, 1899) (status source)
Chlamylla islandica (Odhner, 1937) (new combination reference)
Chlamylla parva (M. Hadfield, 1963) (new combination reference)
Chlamylla polaris (Volodchenko, 1946) (new combination reference)
Corrupta Korshunova, Fletcher & Martynov, 2025 accepted as Coryphella J. E. Gray, 1850 (source of synonymy)
Coryphella J. E. Gray, 1850 (status source)
Coryphella abei Baba, 1987 (status source)
Coryphella alexanderi Ekimova, 2022 (status source)
Coryphella amabilis (Y. Hirano & Kuzirian, 1991) (status source)
Coryphella athadona Bergh, 1875 (status source)
Coryphella borealis Odhner, 1922 (status source)
Coryphella browni Picton, 1980 (status source)
Coryphella californica Bergh, 1904 (status source)
Coryphella candela (Korshunova, Fletcher & Martynov, 2025) (new combination reference)
Coryphella capensis Thiele, 1925 (status source)
Coryphella cerverai (M. A. Fischer, van der Velde & Roubos, 2007) (new combination reference)
Coryphella chriskaugei (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (status source)
Coryphella cooperi T. D. A. Cockerell, 1901 (status source)
Coryphella dana (Millen & Hamann, 2006) (new combination reference)
Coryphella falklandica Eliot, 1907 (status source)
Coryphella fogata (Millen & Hermosillo, 2007) (status source)
Coryphella gracilis (Alder & Hancock, 1844) (status source)
Coryphella insolita (García-Gómez & Cervera, 1989) (status source)
Coryphella lineata (Lovén, 1846) (status source)
Coryphella monicae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (status source)
Coryphella nobilis A. E. Verrill, 1880 (status source)
Coryphella orjani (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (status source)
Coryphella pallida A. E. Verrill, 1900 (status source)
Coryphella sanamyanae (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (status source)
Coryphella trilineata O'Donoghue, 1921 (status source)
Coryphella trophina (Bergh, 1890) (status source)
Coryphellidae Bergh, 1889 accepted as Flabellinidae Bergh, 1889 (source of synonymy)
Coryphellina O'Donoghue, 1929 (status source)
Coryphellina albomarginata (M. C. Miller, 1971) (status source)
Coryphellina arveloi (Ortea & Espinosa, 1998) (additional source)
Coryphellina aurora Ekimova, Deart, Antokhina, Mikhlina & Schepetov, 2022 (status source)
Coryphellina delicata (Gosliner & Willan, 1991) (status source)
Coryphellina exoptata (Gosliner & Willan, 1991) (status source)
Coryphellina flamma Ekimova, Deart, Antokhina, Mikhlina & Schepetov, 2022 (status source)
Coryphellina hamanni (Gosliner, 1994) (status source)
Coryphellina indica (Bergh, 1902) (status source)
Coryphellina lotos Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Coryphellina marcusorum (Gosliner & Kuzirian, 1990) (status source)
Coryphellina pannae Ekimova, Deart, Antokhina, Mikhlina & Schepetov, 2022 (status source)
Coryphellina poenicia (Burn, 1957) (status source)
Coryphellina pseudolotos Ekimova, Deart, Antokhina, Mikhlina & Schepetov, 2022 (status source)
Coryphellina westralis (Burn, 1964) (status source)
Edmundsella Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Edmundsella albomaculata (Pola, Carmona, Calado & Cervera, 2014) (status source)
Edmundsella bertschi (Gosliner & Kuzirian, 1990) (status source)
Edmundsella dushia (Ev. Marcus & Er. Marcus, 1963) (new combination reference)
Edmundsella pedata (Montagu, 1816) (status source)
Edmundsella vansyoci (Gosliner, 1994) (status source)
Eolis rufibranchialis G. Johnston, 1832 accepted as Coryphella verrucosa (M. Sars, 1829) (source of synonymy)
Fjordia Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella J. E. Gray, 1850 (source of synonymy)
Flabellina bandeli (Ev. Marcus, 1976) (basis of record)
Flabellina cavolini (Vérany, 1846) (additional source)
Flabellina enne (Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017) (basis of record)
Flabellina funeka Gosliner & R. J. Griffiths, 1981 (status source)
Flabellina gabinierei (Vicente, 1975) (basis of record)
Flabellina gaditana (Cervera, García-Gómez & F. J. García, 1987) (additional source)
Flabellina ischitana Y. Hirano & T. E. Thompson, 1990 (status source)
Flabellina pellucida (Alder & Hancock, 1843) (status source)
Flabellinidae Bergh, 1889 (status source)
Flabellinopsidae Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Flabellinopsis MacFarland, 1966 (status source)
Flabellinopsis iodinea (J. G. Cooper, 1863) (status source)
Gulenia Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella J. E. Gray, 1850 (source of synonymy)
Kynaria Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Kynaria cynara (Ev. Marcus & Er. Marcus, 1967) (status source)
Launsina Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026 (original description)
Launsina rubropurpurata (Gosliner & Willan, 1991) (new combination reference)
Launsina tanyae Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026 (original description)
Luisella Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 (status source)
Luisella babai (Schmekel, 1972) (status source)
Luisella bulbosa (Ortea & Espinosa, 1998) (new combination reference)
Luisella engeli (Ev. Marcus & Er. Marcus, 1968) (new combination reference)
Luisella llerae (Ortea, 1989) (new combination reference)
Luisella telja (Ev. Marcus & Er. Marcus, 1967) (new combination reference)
Mgueolia Korshunova, Fletcher & Martynov, 2025 (status source)
Mgueolia anastasiae Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026 (original description)
Mgueolia ninae Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026 (original description)
Mgueolia vladimiri Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026 (original description)
Microchlamylla Korshunova, Martynov, Bakken, Evertsen, Fletcher, Mudianta, H. Saito, Lundin, Schrödl & Picton, 2017 accepted as Coryphella J. E. Gray, 1850 (source of synonymy)
Notaeolidia Eliot, 1905 (status source)
Notaeolidia depressa Eliot, 1905 (status source)
Notaeolidia gigas Eliot, 1905 (status source)
Notaeolidia schmekelae Wägele, 1990 (status source)
Sea of Okhotsk for Mgueolia anastasiae Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026
Sea of Okhotsk for Mgueolia ninae Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026
Sea of Okhotsk for Mgueolia vladimiri Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026
Vietnamese part of the South China Sea for Launsina tanyae Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026
Sea of Okhotsk for Mgueolia ninae Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026
Sea of Okhotsk for Mgueolia vladimiri Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026
Vietnamese part of the South China Sea for Launsina tanyae Ekimova, Carmona, Mikhlina, Grishina, Stanovova, Schepetov, Hoover, De Souza-Canal, K. O. Kuznetsov & Á. Valdés, 2026
Etymology
After Launsina (Laun-Sina), the Philippine goddess of the eastern skies, stars and seas who brings in the light of ... [details]
Etymology
Named in honor of Tatiana Antokhina (shortened as Tanya), Russian marine biologist, diver, and the collector of ... [details]
Etymology
Named in honor of Anastasia (also transliterated as Anastassya) Maiorova, leading scientist in National Centre of ... [details]
Etymology
Named in honor of Nina Izotovna Volodchenko, USSR zoologist, the first specialist in systematics and biodiversity ... [details]
Etymology
Named in honor of Vladimir Mordukhovich, leading scientist at the National Centre of Marine Biology, Far East ... [details]
