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Licher, Frank 2000 (1999), Revision der Gattung Typosyllis Langerhans, 1879 (Polychaeta: Syllidae). Morphologie, Taxonomie und Phylogenie. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 551:1-336
23045
Licher, F.
2000
Revision der Gattung Typosyllis Langerhans, 1879 (Polychaeta: Syllidae). Morphologie, Taxonomie und Phylogenie
Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft
551: 1-336
Publication
World Polychaeta Database (WPolyD). Issue date 1999, but published early 2000.
The genus Typosyllis (Langerhans, 1879) (Polychaeta: Syllidae: Syllinae) is revised for the first time. All distinct species of this taxon are described in detail and synonyms listed. Altogether, more than 1850 specimens representing 213 taxa were examined. Twenty-three taxa, previously assigned to Syllis, Langerhansia, Ehlersia or Pionosyllis, are transferred to the genus Typosyllis. Eight species previously assigned to Typosyllis or Syllis are transfered to other syllid taxa. One junior homonym is renamed. The following four species are described as new to science: T. hyllebergi sp. nov., T. mauretanica sp. nov., T. silkeae sp. nov. and T. taiwanensis sp. nov. Three names are treated as Syllinae incertae sedis. Fifty-five names are regarded as younger synonyms of previously described Typosyllis-species. Fifty-seven names are considered nomina dubia. Nine names are considered nomina nuda. The names Syllis atlantico and S. atlantica lineata originally erected as nomina nova have to be eliminated. According to this revision the genus Typosyllis comprises 107 distinct species. A key for identification of all species is provided. Four lectotypes and several paralectotypes are designated: Typosyllis vittata (Grube, 1840) (MPW 673), T. valida Grube, 1857 (ZMUC 881; ZMUC 890), T. neglecta (Grube, 1870) comb. nov. (ZMB Verm. 843 (LT); ZMB Verm. 11074 (PLT)) und T. violacea (Grube, 1870) comb. nov. (ZMB Verm. 844; ZMB 11073). For two species neotypes are designated: Typosyllis cornuta (Rathke, 1843) (NTNU) und T. prolifera (Krohn, 1852) (NHMG Inve-21334). In the computer-based, cladistic analysis of Typosyllis revealed 3 large monophyletic groups of complexes: the complex-group cornuta (22 species, 6 species-groups), the complex-group prolifera (49 species, 8 species-groups) and the complex-group armillaris (25 species, 4 species-groups). Eleven species cannot be assigned with certainty to any of these 18 species-groups. It is shown that the taxa previously assigned to the genus Langerhansia (species with falcigerous and pseudospinigerous chaetae) probably form a monophylum, whereas the taxa previously assigned to the genus Typosyllis (species with falcigerous chaetae only) form a paraphylum. Nevertheless, the taxa Langerhansia is eliminated by referring the respective species to the genus Typosyllis, since the distinction of these two groups is based only on slight, gradual differences in some characters, providing no sound reasoning for a correct assignment of some species to Typosyllis or to Langerhansia.
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Branchiosyllis bathyalis (Kirkegaard, 1995) (basis of record)
Ehlersia cornuta (Rathke, 1843) accepted as Syllis cornuta Rathke, 1843 (source of synonymy)
Ehlersia ferrugina Langerhans, 1881 accepted as Paraehlersia ferrugina (Langerhans, 1881) (source of synonymy)
Haplosyllis cephalata Verrill, 1900 (additional source)
Haplosyllis spongicola (Grube, 1855) (additional source)
Haplosyllis spongiphila (Verrill, 1885) (basis of record)
Langerhansia Czerniavsky, 1881 accepted as Syllis Lamarck, 1818 (source of synonymy)
Langerhansia cornuta (Rathke, 1843) accepted as Syllis cornuta Rathke, 1843 (source of synonymy)
Syllis (Ehlersia) cornuta Rathke, 1843 accepted as Syllis cornuta Rathke, 1843 (source of synonymy)
Syllis (Ehlersia) ferrugina (Langerhans, 1881) accepted as Paraehlersia ferrugina (Langerhans, 1881) (source of synonymy)
Syllis (Ehlersia) heterochaeta Moore, 1909 accepted as Syllis heterochaeta Moore, 1909 (new combination reference)
Syllis (Ehlersia) hyperioni Dorsey & Phillips, 1987 accepted as Syllis hyperioni Dorsey & Phillips, 1987 (source of synonymy)
Syllis (Haplosyllis) Langerhans, 1879 accepted as Haplosyllis Langerhans, 1879 (source of synonymy)
Syllis (Haplosyllis) spongicola Grube, 1855 accepted as Haplosyllis spongicola (Grube, 1855) (source of synonymy)
Syllis (Langerhansia) Czerniavsky, 1881 accepted as Syllis Lamarck, 1818 (source of synonymy)
Syllis (Langerhansia) cornuta Rathke, 1843 accepted as Syllis cornuta Rathke, 1843 (source of synonymy)
Syllis (Langerhansia) ferrugina (Langerhans, 1881) accepted as Paraehlersia ferrugina (Langerhans, 1881) (source of synonymy)
Syllis (Typosyllis) armillaris (O.F. Müller, 1776) accepted as Syllis armillaris (O.F. Müller, 1776) (source of synonymy)
Syllis (Typosyllis) gerlachi (Hartmann-Schröder, 1960) accepted as Syllis gerlachi (Hartmann-Schröder, 1960) (source of synonymy)
Syllis (Typosyllis) hyalina Grube, 1863 accepted as Syllis hyalina Grube, 1863 (source of synonymy)
Syllis (Typosyllis) variegata Grube, 1860 accepted as Syllis variegata Grube, 1860 (source of synonymy)
Syllis abyssicola Ehlers, 1874 (status source)
Syllis alternata Moore, 1908 (redescription)
Syllis amica Quatrefages, 1866 (additional source)
Syllis armillaris (O.F. Müller, 1776) (redescription)
Syllis brevicollis Ehlers, 1874 (status source)
Syllis closterobranchia Schmarda, 1861 accepted as Syllis armillaris (O.F. Müller, 1776) (source of synonymy)
Syllis cornuta Rathke, 1843 (redescription)
Syllis ferrugina (Langerhans, 1881) accepted as Paraehlersia ferrugina (Langerhans, 1881) (source of synonymy)
Syllis ferruginea (Langerhans, 1881) accepted as Paraehlersia ferrugina (Langerhans, 1881) (source of synonymy)
Syllis gerlachi (Hartmann-Schröder, 1960) (redescription)
Syllis hyalina Grube, 1863 (redescription)
Syllis prolifera Krohn, 1852 (redescription)
Syllis setubalensis McIntosh, 1885 accepted as Haplosyllis spongicola (Grube, 1855) (source of synonymy)
Syllis spongicola Grube, 1855 accepted as Haplosyllis spongicola (Grube, 1855) (source of synonymy)
Syllis spongiphila Verrill, 1885 accepted as Haplosyllis spongiphila (Verrill, 1885) (new combination reference)
Syllis tristanensis Day, 1954 accepted as Syllis hyalina Grube, 1863 (source of synonymy)
Syllis valida Grube, 1857 (source of synonymy)
Syllis variegata Grube, 1860 (redescription)
Syllis vittata Grube, 1840 (redescription)
Trypanosyllis gigantea (McIntosh, 1885) accepted as Trypanedenta gigantea (McIntosh, 1885) (additional source)
Typosyllis Langerhans, 1879 accepted as Syllis Lamarck, 1818 (source of synonymy)
Typosyllis Langerhans, 1879 accepted as Syllis Lamarck, 1818 (status source)
Typosyllis (Typosyllis) armillaris (O.F. Müller, 1776) accepted as Syllis armillaris (O.F. Müller, 1776) (source of synonymy)
Typosyllis (Typosyllis) hyalina (Grube, 1863) accepted as Syllis hyalina Grube, 1863 (source of synonymy)
Typosyllis armillaris (O.F. Müller, 1776) accepted as Syllis armillaris (O.F. Müller, 1776) (basis of record)
Typosyllis bathyalis Kirkegaard, 1995 accepted as Branchiosyllis bathyalis (Kirkegaard, 1995) (new combination reference)
Typosyllis brachychaeta (Schmarda, 1861) accepted as Syllis armillaris (O.F. Müller, 1776) (source of synonymy)
Typosyllis cornuta (Rathke, 1843) accepted as Syllis cornuta Rathke, 1843 (source of synonymy)
Typosyllis cornuta (Rathke, 1843) accepted as Syllis cornuta Rathke, 1843 (basis of record)
Typosyllis hyalina (Grube, 1863) accepted as Syllis hyalina Grube, 1863 (basis of record)
Typosyllis parateinopterona Licher, 1999 (original description)
Typosyllis variegata (Grube, 1860) accepted as Syllis variegata Grube, 1860 (basis of record)
Typosyllis vittata (Grube, 1840) accepted as Syllis vittata Grube, 1840 (additional source)
Holotype AHF Poly 888, geounit North West Atlantic, identified as Typosyllis parateinopterona Licher, 1999
 Depth range
 for Syllis cerina Grube, 1878

11-16 m. [details]
 Depth range
 for Typosyllis crassicirrata Treadwell, 1925

Shallow water and 53-115 m.  [details]
 Depth range
 for Typosyllis garciai (Campoy, 1982)

1-35 m. [details]
 Depth range
 for Typosyllis hawaiiensis Hartmann-Schröder, 1965

Eulittoral.  [details]
 Depth range
 for Typosyllis hyllebergi Licher, 1999

Eulittoral.  [details]
 Depth range
 for Typosyllis lutea Hartmann-Schröder, 1960

Eulittoral to sublittoral. Between 2-3 m at the type locality.  [details]
 Depth range
 for Typosyllis mauretanica Licher, 1999

Eulittoral.  [details]
 Depth range
 for Typosyllis parateinopterona Licher, 1999

Continental shelf and shallow to deep slope depths, 97-2900 m. [details]
 Depth range
 for Typosyllis silkeae Licher, 1999

Eulittoral.  [details]
 Distribution
 for Branchiosyllis verruculosa (Augener, 1913)

Australia: Queensland; South Australia; Western Australia (Shark Bay and Albany District). Indonesia. Eastern Africa: ... [details]
 Distribution
 for Syllis cerina Grube, 1878

Philippine Islands (Bohol); Western Australia (Shark Bay).  [details]
 Distribution
 for Typosyllis anops (Ehlers, 1897)

South Africa; Argentina; Chile; Antarctic Ocean; Kerguelen; New Zealand; Australia; South China.  [details]
 Distribution
 for Typosyllis crassicirrata Treadwell, 1925

Pacific Ocean: Hawaiian Islands; Polynesia; NW Australia. [details]
 Distribution
 for Typosyllis garciai (Campoy, 1982)

Mediterranean Sea. Caribbean Sea: Cuba.  [details]
 Distribution
 for Typosyllis gerlachi Hartmann-Schröder, 1960

Red Sea; Mediterranean Sea; Indian Ocean; Caribbean Sea; Gulf of Mexico.  [details]
 Distribution
 for Typosyllis hawaiiensis Hartmann-Schröder, 1965

Pacific Ocean: Hawaiian Islands.  [details]
 Distribution
 for Typosyllis hyllebergi Licher, 1999

Red Sea, Gulf of Aqaba and Suez Canal.  [details]
 Distribution
 for Typosyllis lutea Hartmann-Schröder, 1960

Red Sea: Saudi Arabia (Sarso Island). Circumtropical to subtropical.  [details]
 Distribution
 for Typosyllis mauretanica Licher, 1999

Mauritania, Banc d'Arguin.  [details]
 Distribution
 for Typosyllis parateinopterona Licher, 1999

Atlantic Ocean: continental slope off New England (USA); continental shelves of Guiana and Brazil (off Pernambuco). [details]
 Distribution
 for Typosyllis schulzi Hartmann-Schröder, 1960

Red Sea: Egypt (Hurghada). Indian Ocean: Mozambique (Xai Xai). Mediterranean Sea.  [details]
 Distribution
 for Typosyllis silkeae Licher, 1999

Taiwan.  [details]
 Distribution
 for Typosyllis truncata (Haswell, 1920)

See note for Syllis truncata [details]
 Editor's comment
 for Syllis brevis Schmarda, 1861

The description of the species Syllis brevis Schmarda, 1861 seems to indicate a specimen belonging to the genus ... [details]
 Editor's comment
 for Syllis singulisetis Grube, 1878

Licher (1999: 307-308) revised the type material deposited at the MPW, considering it to be a nomen dubium due to ... [details]
 Editor's comment
 for Syllis tristanensis Day, 1954

Day (1954) did not state explicitly which of the 46 stations where the new species was collected was the type ... [details]
 Etymology
 for Syllis nigricirris Grube, 1863

The specific epithet nigricirris refers to the dorsal black cirri that characterizes the species.  [details]
 Etymology
 for Typosyllis hawaiiensis Hartmann-Schröder, 1965

The specific epithet hawaiiensis refers to the type locality of the species, the Hawaiian Islands.  [details]
 Etymology
 for Typosyllis hyllebergi Licher, 1999

The species is dedicated to Dr. Jørgen Hylleberg, from the University of Århus, Denmark, who collected the type ... [details]
 Etymology
 for Typosyllis lutea Hartmann-Schröder, 1960

The specific epithet lutea, feminine form of the Latin word luteus, meaning 'yellowish' or 'sallow', refers to the ... [details]
 Etymology
 for Typosyllis mauretanica Licher, 1999

The specific epithet mauretanica refers to the type locality of the species, Mauritania.  [details]
 Etymology
 for Typosyllis parateinopterona Licher, 1999

The specific epithet parteinopterona refers to the great length of the blades, being composed by the old Greek ... [details]
 Etymology
 for Typosyllis silkeae Licher, 1999

The species is dedicated to the wife of the author, Silke.  [details]
 Habitat
 for Typosyllis garciai (Campoy, 1982)

On tubes of sabellids and among green algae, eulittoral. [details]
 Habitat
 for Typosyllis hawaiiensis Hartmann-Schröder, 1965

Among eulittoral algae.  [details]
 Habitat
 for Typosyllis hyllebergi Licher, 1999

Eulittoral.  [details]
 Habitat
 for Typosyllis mauretanica Licher, 1999

AEulittoral, among seagrasses.  [details]
 Habitat
 for Typosyllis parateinopterona Licher, 1999

Continental shelf and slope. Further information lacking.  [details]
 Habitat
 for Typosyllis silkeae Licher, 1999

Eulittoral, in sand with coral rubble.  [details]
 Homonymy
 for Langerhansia anoculata Hartman & Fauchald, 1971

With the move of the species to the genus Typosyllis performed by Licher (1999) the species became a secondary ... [details]
 Homonymy
 for Typosyllis parateinopterona Licher, 1999

New name for Langerhansia anoculata Hartman & Fauchald, 1971 nec Hartmann-Schroder, 1962. With the move of ... [details]
 Neotype
 for Syllis (Ehlersia) heterochaeta Moore, 1909

Dorsey & Phillips (1987) searched unsuccessfully for the holotype of Syllis (Ehlersia) heterochaeta at the ... [details]
 Nomenclature
 for Langerhansia (Syllis) [lapsus for Syllis (Langerhansia)]

An impossible apparent reversed-presentation of subgenus Langerhansia by Czerniavsky, 1881. It is clear that while ... [details]
 Nomenclature
 for Syllis gracilis Schmarda, 1861

Licher (1999: 54-57) revised the type material of Syllis gracilis Schmarda, 1861, and determined that it was a ... [details]
 Nomenclature
 for Syllis krohnii Ehlers, 1864

Licher (1999: 205, footnote) corrects the specific epithet from krohnii to krohni, evoking article 31.1.2. of the ... [details]
 Nomenclature
 for Typosyllis (Typosyllis) luteoides hawaiensis Hartmann-Schröder, 1965

The specific epithet was corrected by Licher (1999: 131) from hawaiensis to hawaiiensis.  [details]
 Nomenclature
 for Typosyllis anops (Ehlers, 1897)

Licher (1999: 54-57) revised the type material of Syllis gracilis Schmarda, 1861, and determined that it was a ... [details]
 Nomenclature
 for Typosyllis hawaiiensis Hartmann-Schröder, 1965

The specific epithet was corrected by Licher (1999: 131) from the original hawaiensis to the current hawaiiensis.  [details]
 Nomenclature
 for Typosyllis heterocirra Hartmann-Schröder, 1960

According to Licher (1999: 264, footnote) the correct latinization of the specific epithet is heterocirrata, ... [details]
 Nomenclature
 for Typosyllis heterocirrata Hartmann-Schröder, 1960

According to Licher (1999: 264, footnote) the correct latinization of the specific epithet is heterocirrata, ... [details]
 Status
 for Gnathosyllis diplodonta Schmarda, 1861

Hartman (1959: 225) considered that the species belonged to the genus Syllis, being however indeterminable to ... [details]
 Status
 for Syllis abyssicola Ehlers, 1874

The species was considered to be a nomen dubium by Licher (1999: 304).  [details]
 Status
 for Syllis attenuata Knox, 1960

Licher 2000:297 placed as species inquirenda in Typosyllis (now disused), possibly because the type could not be found. [details]
 Status
 for Syllis brevicollis Ehlers, 1874

The species was considered to be a nomen dubium by Licher (1999: 304).  [details]
 Synonymy
 for Syllis sardai San Martín, 1992

The species was synonymized with Syllis variegata Grube, 1860 (as Typosyllis) by Licher (1999: 108).  [details]
 Synonymy
 for Typosyllis Langerhans, 1879

Licher (2000), in a monumental and painstaking monograph on genus Typosyllis, maintained the genus named by ... [details]
 Taxonomic remark
 for Syllis ortizi San Martín, 1992

According to Licher et al. (1995) and Licher (1999), the paratypes USNM 75308 and USNM 75312 of Syllis ortizi San ... [details]
 Taxonomic remark
 for Typosyllis magnipectinis (Storch, 1967)

According to Licher et al. (1995) and Licher (1999), the paratypes USNM 75308 and USNM 75312 of Syllis ortizi San ... [details]
 Taxonomy
 for Branchiosyllis cirropunctata (Michel, 1909)

Licher (1999) and San Martín (2003) erroneously considered Syllis cirropunctata as a synonym of Branchiosyllis ... [details]
 Taxonomy
 for Chaetosyllis oerstedi Malmgren, 1867

Current taxon was considered as a junior synonym of Syllis cornuta Rathke, 1843 by Carus (1885: 228), Soulier ... [details]
 Taxonomy
 for Haplosyllis cephalata Verrill, 1900

Current taxon was generally considered as a junior synonym of Haplosyllis spongicola (Grube, 1855), but was ... [details]
 Taxonomy
 for Pionosyllis aciculigrossa San Martín, 1990

Moved to genus Typosyllis by Licher (1999), and Syllis in Amaral & Rossi-Wongtschowski (2004).  [details]
 Taxonomy
 for Pionosyllis lucida Chamberlin, 1919

Moved to genus Typosyllis by Hartman (1959: 217, 234). Considered as a nomen dubium by Licher (1999: 304)  [details]
 Taxonomy
 for Syllis (Typosyllis) augeneri Haswell, 1920

The species was moved to genus Typosyllis Langerhans, 1879 by Hartman (1959: 230, 234), in spite of the fact that ... [details]
 Taxonomy
 for Syllis (Typosyllis) grandigularis Verrill, 1900

San Martín (1992) first and Licher (1999) later revised the type material and considered the species to be synonym ... [details]
 Taxonomy
 for Syllis (Typosyllis) punctulata Haswell, 1920

Moved to genus Typosyllis by Hartman (1959: 235).
Licher (1999: 307) considered the species as nomen dubium, due ... [details]
 Taxonomy
 for Syllis brevis Schmarda, 1861

In the absence of type material Licher (1999: 304) considered the species to be a nomen dubium.  [details]
 Taxonomy
 for Syllis erythropis Grube, 1878

Licher (1999) considered the species as a nomen dubium, as the type material did not enable a redescription.  [details]
 Taxonomy
 for Syllis fusicornis Schmarda, 1861

Licher (1999: 305) in the absence of type material considered the species to be a nomen dubium.  [details]
 Taxonomy
 for Syllis grandigularis Verrill, 1900

San Martín (1992) first and Licher (1999) later revised the type material and considered the species to be synonym ... [details]
 Taxonomy
 for Syllis macroura Schmarda, 1861

Hartman (1959: 227), considered Syllis macroura as being possibly Opisthosyllis australis Augener, 1913. In the ... [details]
 Taxonomy
 for Syllis magdalena Wesenberg-Lund, 1962

Moved to genus Typosyllis by Licher (1999) but considered nowadays to belong to Syllis.  [details]
 Taxonomy
 for Syllis normannica Claparède, 1863

Current taxon junior synonym of species listed. According to Licher (1999: 306) the species should be considered as ... [details]
 Taxonomy
 for Syllis remex Chamberlin, 1919

According to Licher (1999: 307) the species is not a syllid, being the holotype an epitoke form with hypertrophied ... [details]
 Taxonomy
 for Syllis singulisetis Grube, 1878

Licher (1999) considered the species as nomen dubium. The holotype is in poor condition, with the soft tissues ... [details]
 Taxonomy
 for Syllis solida Grube, 1878

Species uncertain. Augener (1913) stated that Syllis solida was very similar to Branchiosyllis exilis (Gravier, ... [details]
 Taxonomy
 for Thoe fusiformis Kinberg, 1866

The species was considered as a possible synonym of Syllis variegata Grube, 1860 by Augener (1913: 190-192; 1922: ... [details]
 Type locality
 for Syllis nigrescens Grube, 1878

Philippine Islands. Grube (1878: VII) comments that probably Syllis nigrescens was collected in Singapore, but ... [details]
 Type locality
 for Typosyllis hawaiiensis Hartmann-Schröder, 1965

Pacific Ocean: Tern Island, French Frigate Shoals (Northwestern Hawaiian Islands).  [details]
 Type locality
 for Typosyllis hyllebergi Licher, 1999

Egypt (marine), Dahab, El Kura Bay (geocoordinates not provided, but estimated with gazetteer to be approximately ... [details]
 Type locality
 for Typosyllis mauretanica Licher, 1999

Mauritania (marine), Banc d' Arguin, about 100 km south of the city of Nouadhibou (geocoordinates not provided, but ... [details]
 Type locality
 for Typosyllis nipponica Imajima, 1966

Pacific Ocean, Japan. The species was described with base on specimens from several different localities in Japan. ... [details]
 Type locality
 for Typosyllis parateinopterona Licher, 1999

Atlantic Ocean, northwestern section, deep slope off New England, Atlantic coast of the USA (39º43.6'N, 70º37.4'W).  [details]
 Type locality
 for Typosyllis silkeae Licher, 1999

Taiwan (marine), Liu-chiu-yu (22º20'30"N, 120º22'00"E).  [details]
 Type material
 for Chaetosyllis oerstedi Malmgren, 1867

The type material was not studied or located by Licher (1999: 300).  [details]
 Type material
 for Ehlersia (Syllis) rosea Langerhans, 1879

The type material was not located by Licher (1999: 45). However, the NHM, Wien, holds 6 specimens (NHMW 668) ... [details]
 Type material
 for Gnathosyllis diplodonta Schmarda, 1861

Not found by Licher (1999), probably lost.  [details]
 Type material
 for Syllis (Typosyllis) plessisi Rullier, 1972

The type material includes only the holotype, mounted in a slide. It was initially deposited at the collections of ... [details]
 Type material
 for Syllis abyssicola Ehlers, 1874

Type material unavailable, probably lost, not located by Licher (1999: 304). [details]
 Type material
 for Syllis brevicollis Ehlers, 1874

The type material was not found by Licher (1999: 304).  [details]
 Type material
 for Syllis brevis Schmarda, 1861

Type material not found by Licher (1999: 304), who considered it to be probably lost.  [details]
 Type material
 for Syllis erythropis Grube, 1878

Holotype deposited at the MPW (No. 393). The specimen dried (Wiktor, 1980), and Licher (1999) considered the ... [details]
 Type material
 for Syllis flaccida Grube, 1878

Grube (1878) refers the collection of 4 specimens from Aibuhit (maybe Aimellik?; Palau Islands) and Camiguin ... [details]
 Type material
 for Syllis fusicornis Schmarda, 1861

Type material not found by Licher (1999: 305), who considered it to be probably lost. [details]
 Type material
 for Syllis hyalina Grube, 1863

The species was described with base on specimens from 3 different locations: Veli Lošinj (Lussin Grande), Nerezine ... [details]
 Type material
 for Syllis inflata Marenzeller, 1879

Syntypes (2) deposited at the Naturhistorisches Museum, Wien, Austria: NHMW 618.  [details]
 Type material
 for Syllis lycochaetus Grube, 1878

Licher (1999) did not find the type material, it might have been lost.  [details]
 Type material
 for Syllis macroura Schmarda, 1861

Type material not found by Licher (1999: 306).  [details]
 Type material
 for Syllis nigrescens Grube, 1878

Holotype deposited at the MPW (No. 397), in two fragments.  [details]
 Type material
 for Syllis ortizi San Martín, 1992

Paratypes USNM 75308 and USNM 75312 of Syllis ortizi San Martín, 1992 were considered as being Typosyllis ... [details]
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