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Brodsky, A.L. (1929). Фауна водоемов пустыни Кара-кум - The fauna of the basins of the desert Kara-Kum. Acta Universitatis Asiae Mediae (XIIa, Geogr.). 5:1-42, figs. 1-9, pls. 1-2, tabs. 1-3, map. (Russian with English summary.).
Brodsky, A.L. (1929). Фауна водоемов пустыни Кара-кум - The fauna of the basins of the desert Kara-Kum. Acta Universitatis Asiae Mediae (XIIa, Geogr.). 5:1-42, figs. 1-9, pls. 1-2, tabs. 1-3, map. (Russian with English summary.).