A glypticid with very strongly developed, coarse epistroma over its aboral surface, and with rudimentary aboral interambulacral tubercles that are hardly distinguishable from amongst the epistroma.
Smith, A.B. (2016). British Jurassic Regular Echinoids. <em>Monograph of the Palaeontographical Society.</em> 170 (646): 69-176, pls 42-82.