Cretaceous-Eocene [details]
Ilerdian, Early Eocene [details]
exact locality unknown, near Czarna, Wiselka, Silesia, Poland [details]
Zollhaus Quarry, ca. 25 km W of Lake Thun, Prealps, Switzerland [details]
HT: BMNH T762 [details]
HT: BMNH T763 (middle of three traces); PT: BMNH T763 (outer two traces) [details]