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Clark, A. H., Clark, A. M. (1967). A monograph of the existing crinoids. Bulletin of the United States National Museum. (82): 1-860.
133557
Clark, A. H., Clark, A. M.
1967
A monograph of the existing crinoids
Bulletin of the United States National Museum
(82): 1-860
Publication
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BibTex (BibDesk, LaTeX)
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2013-01-12 18:30:12Z
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2019-07-26 13:52:22Z
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Anisometra John, 1939 (additional source)
Anisometra frigida John, 1939 (basis of record)
Antedon de Fréminville, 1811 (additional source)
Antedon abyssorum Carpenter, 1888 accepted as Thaumatometra abyssorum (Carpenter, 1888) (basis of record)
Antedon erythrizon AH Clark, 1907 accepted as Psathyrometra fragilis (AH Clark, 1907) (additional source)
Antedon fragilis AH Clark, 1907 accepted as Psathyrometra fragilis (AH Clark, 1907) (basis of record)
Antedon remota Carpenter, 1888 accepted as Tonrometra remota (Carpenter, 1888) (basis of record)
Antedonidae Norman, 1865 (basis of record)
Antedoninae (Norman, 1865) (additional source)
Anthometrina adriani (Bell, 1908) (additional source)
Atelecrinidae Bather, 1899 (additional source)
Atelecrinus Carpenter, 1881 (additional source)
Atelecrinus balanoides Carpenter, 1881 (additional source)
Atelecrinus helgae AH Clark, 1913 (additional source)
Bathymetrinae AH Clark, 1909 (basis of record)
Caryometra AH Clark, 1936 (basis of record)
Caryometra AH Clark, 1936 (additional source)
Caryometra alope AH Clark, 1940 (basis of record)
Caryometra lisa AH Clark, 1940 (basis of record)
Coccometra AH Clark, 1908 (basis of record)
Coccometra guttata AH Clark, 1918 (basis of record)
Coccometra hagenii (Pourtalès, 1868) (basis of record)
Coccometra nigrolineata AH Clark, 1918 (basis of record)
Eometra Clark, 1936 (additional source)
Eometra antarctica (AH Clark, 1915) (basis of record)
Eometra weddelli John, 1939 (in Vaney & John, 1939) (basis of record)
Eumorphometra Clark, 1915 (additional source)
Eumorphometra fraseri John, 1938 (basis of record)
Eumorphometra hirsuta (Carpenter, 1888) (basis of record)
Eumorphometra marri John, 1938 (basis of record)
Florometra AH Clark, 1913 (additional source)
Florometra mawsoni AH Clark, 1937 accepted as Promachocrinus mawsoni (AH Clark, 1937) (basis of record)
Hathrometra sarsii (Düben & Koren, 1846) accepted as Hathrometra tenella (Retzius, 1783) (basis of record)
Hathrometra tenella (Retzius, 1783) (basis of record)
Heliometra AH Clark, 1907 (basis of record)
Heliometra glacialis (Owen, 1833 ex Leach MS) (basis of record)
Heliometrinae AH Clark, 1909 (basis of record)
Isometra AH Clark, 1908 (basis of record)
Isometra flavescens John, 1938 (basis of record)
Isometra graminea John, 1938 (additional source)
Isometra hordea John, 1938 (additional source)
Isometra vivipara Mortensen, 1917 (additional source)
Kempometra John, 1938 (additional source)
Kempometra grisea John, 1938 (basis of record)
Leptometra AH Clark, 1908 (additional source)
Leptometra celtica (M'Andrew & Barrett, 1857) (basis of record)
Leptometra phalangium (Müller, 1841) (basis of record)
Nanometra AH Clark, 1907 (basis of record)
Nanometra clymene AH Clark, 1912 (basis of record)
Notocrinidae Mortensen, 1918 (basis of record)
Notocrinus Mortensen, 1917 (basis of record)
Notocrinus mortenseni John, 1938 (basis of record)
Notocrinus virilis Mortensen, 1917 (basis of record)
Orthometra hibernica (AH Clark, 1913) (basis of record)
Pentametrocrinidae AH Clark, 1908 (basis of record)
Pentametrocrinus AH Clark, 1908 (additional source)
Pentametrocrinus atlanticus (Perrier, 1883) (additional source)
Pentametrocrinus tuberculatus (AH Clark, 1907) (basis of record)
Pentametrocrinus varians (Carpenter, 1882) (additional source)
Phrixometra Clark, 1921 (additional source)
Phrixometra exigua (Carpenter, 1888) (new combination reference)
Phrixometra longipinna (Carpenter, 1888) (basis of record)
Phrixometra longipinna antarctica John, 1938 (basis of record)
Phrixometra rayneri John, 1938 (basis of record)
Poliometra AH Clark, 1923 (additional source)
Poliometra prolixa (Sladen, 1881) (basis of record)
Promachocrinus Carpenter, 1879 (basis of record)
Promachocrinus kerguelensis Carpenter, 1879 (basis of record)
Promachocrinus vanhoffenianus Minckert, 1905 (basis of record)
Psathyrometra AH Clark, 1907 (additional source)
Psathyrometra bigradata (Hartlaub, 1895) (additional source)
Psathyrometra borealis AH Clark, 1908 accepted as Psathyrometra fragilis (AH Clark, 1907) (basis of record)
Psathyrometra congesta AH Clark, 1908 (additional source)
Psathyrometra erythrizon (AH Clark, 1907) accepted as Psathyrometra fragilis (AH Clark, 1907) (basis of record)
Psathyrometra fragilis (AH Clark, 1907) (additional source)
Psathyrometra mira AH Clark, 1909 accepted as Athrypsometra mira (AH Clark, 1909) (basis of record)
Psathyrometra profundorum AH Clark, 1908 accepted as Psathyrometra fragilis (AH Clark, 1907) (basis of record)
Sarametra AH Clark, 1917 (additional source)
Sarametra triserialis (AH Clark, 1908) (additional source)
Solanometra Clark, 1911 (basis of record)
Solanometra antarctica (Carpenter, 1881) (basis of record)
Thaumatocrinus Carpenter, 1883 (additional source)
Thaumatocrinus investigatoris AH Clark in AH Clark & AM Clark, 1967 (original description)
Thaumatocrinus jungerseni AH Clark, 1915 (additional source)
Thaumatocrinus renovatus Carpenter, 1884 (additional source)
Thaumatocrinus rugosus (AH Clark, 1908) (basis of record)
Thaumatometra abyssorum (Carpenter, 1888) (basis of record)
Thaumatometra alternata (Carpenter, 1888) (basis of record)
Thaumatometra septentrionalis AH Clark, 1918 (basis of record)
Thysanometrinae AH Clark, 1909 (basis of record)
Tonrometra AH Clark, 1917 (additional source)
Tonrometra multicirra AH Clark, 1929 (basis of record)
Tonrometra remota (Carpenter, 1888) (basis of record)
Tonrometra spinulifera (John, 1939) (new combination reference)
Trichometra AH Clark, 1908 (additional source)
Trichometra aspera AH Clark, 1908 accepted as Trichometra cubensis (Pourtalès, 1869) (basis of record)
Trichometra cubensis (Pourtalès, 1869) (basis of record)
Trichometra delicata AH Clark, 1911 (basis of record)
Trichometra hibernica AH Clark, 1913 accepted as Orthometra hibernica (AH Clark, 1913) (basis of record)
Zenometra AH Clark, 1907 (additional source)
Holotype IMC, identified as Thaumatocrinus investigatoris AH Clark in AH Clark & AM Clark, 1967
Holotype NRS (Antarctic sta. 58), identified as Isometra vivipara Mortensen, 1917
Syntype BMNH (Discovery Investigations sta. 170), identified as Isometra hordea John, 1938
Syntype BMNH (Discovery Investigations sta. 190,1652,1872), identified as Isometra graminea John, 1938
Holotype USNM 22674, identified as Coccometra nigrolineata AH Clark, 1918
Holotype USNM 22675, identified as Coccometra guttata AH Clark, 1918
Holotype UMZC MCZ 7, identified as Coccometra hagenii (Pourtalès, 1868)
 Diagnosis

Andrometra with centrodorsal conical, as high as broad at the base; cirri 50. Second and third pinnules with 15 ... [details]

 Diagnosis

Andrometra with centrodorsal low hemispherical to almost columnar; cirri not more than 35. Second pinnule with ... [details]

 Diagnosis

Annametra with P1 and P2 with ~18 segments. Arms ~30 mm and cirri ~5 mm long. [details]

 Diagnosis

Annametra with P1 and P2 of up to 35 very short segments. Arms ~50 mm long in adult. Longest cirri up to 15 mm long. [details]

 Diagnosis

Antedon with centrodorsal flattened hemispherical. Cirri ~30, in three crowded marginal rows, of 10-12 cirrals; ... [details]

 Diagnosis

Antedon bifida with P2 similar to and same length as P3 but with no gonad. P1 about twice as long as following two ... [details]

 Diagnosis

Antedon bifida with distal cirrals strongly compressed laterally, up to twice as wide in lateral view as proximal ... [details]

 Diagnosis

Antedon with not more than 35 cirri, of not over 17 (rarely more than 15) cirrals; distal cirrals strongly ... [details]

 Diagnosis

Antedon with arms to ~50 mm long. Cirri to 40, of 16-18 cirrals; length to ~25% arm length; longer proximal cirrals ... [details]

 Diagnosis

Antedon incommoda with a relatively smaller aboral pole (0.4-0.9 mm across) with center sometimes very slightly ... [details]

 Diagnosis

Antedon incommoda with centrodorsal discoidal, with aboral pole quite flat although variable in width and with ... [details]

 Diagnosis

Antedon with cirri ~30, of 10-12 cirrals, short (about a fifth the arm length), rather stout, strongly recurved ... [details]

 Diagnosis

Small Antedon with arms up to 40 mm long; cirri 10-20,of 12-17 cirrals, long, (more than a fourth the arm length), ... [details]

 Diagnosis

Antedon with centrodorsal hemispherical, more or less flattened, with fairly large convex polar area. Cirri 15-30, ... [details]

 Diagnosis

Antedon with 18-30 (usually 20-23) cirrals; all but first 2-3 are longer than wide and centrally constricted; distal ... [details]

 Diagnosis

Antedon with P2 stouter basally and two-thirds the length of P1, which has 18-20 segments. Cirri ~30, of up to 11 ... [details]

 Diagnosis

Small Antedon with arms to only 35 mm (rarely 20 mm) long. Cirri <40, about a fifth as long as arms, slender, delicate, of 8-11 greatly elongated segments with greatly expanded distal ends (weker distally); longest with L/W ratio ~4; antepenultimate with L/W ratio ~2; penultimate with L/W ratio 1.3-1.5; opposing spine prominent. P2 is about half as long as P1. Centrodorsal ... [details]

 Diagnosis

Antedon with P2 intermediate in length between P1 and P3, and resembling P1. Cirri usually 40, of at most 18 ... [details]

 Diagnosis

Antedon with arms up to 70 mm long. Centrodorsal flattened hemispherical. Cirri in largest specimens about ... [details]

 Diagnosis

Argyrometra with first three pinnules successively longer; pinnulars with rather prominently everted and spinous ... [details]

 Diagnosis

Argyrometra with first three pinnules successively longer. Cirri of 12-17 cirrals. Holotype with arms ~30 mm long. [details]

 Diagnosis

Athrypsometra with low conical centrodorsal 1.7 mm across base and 1.0 mm tall. Radial areas each with 2 columns of ... [details]

 Diagnosis

Athrypsometra with centrodorsal sharply conical, ~1.8 times wider across base than tall; sockets in 2 (apical) to 4 ... [details]

 Diagnosis

Athrypsometra with centrodorsal 1.9 mm across base and 1.6 mm tall, with 4-5 cirrus sockets in each radial area in ... [details]

 Diagnosis

Athrypsometra with centrodorsal almost as tall as basal diameter in large specimens but relatively lower in small ... [details]

 Diagnosis

Atopocrinus with cirrus sockets in 10 columns, two per radial area. [details]

 Diagnosis

Same as genus. [details]

 Diagnosis

Bathymetra with 10-15 cirri around aboral pole, directed aborally; none around centrodorsal base. Radials form ... [details]

 Diagnosis

Bathymetra with centrodorsal having flattened aboral pole with cirrus sockets alternating irregularly around the ... [details]

 Diagnosis

Same as genus. [details]

 Diagnosis

Caryometra with centrodorsal about 1.25 times taller than wide. Cirri in 15 crowded columns, slender, of moderate ... [details]

 Diagnosis

Caryometra with centrodorsal low conical, wider than tall. Cirri in 10 columns, long, very slender, scarcely curved ... [details]

 Diagnosis

Caryometra with centrodorsal about as tall as wide; aboral apex papillose. Cirri in 15 columns, long, very slender, ... [details]

 Diagnosis

Caryometra with centrodorsal rounded conical, about as tall as wide. Cirri in 10 columns, short, strongly curved ... [details]

 Diagnosis

Caryometra with centrodorsal slightly taller than wide. Cirri in 15 columns, long, slender, of 27-34 cirrals; ... [details]

 Diagnosis

Caryometra with centrodorsal sharply conical, up to 1.5 times taller than wide at the base. Cirri in 15 columns ... [details]

 Diagnosis

Cyclometra with cirri ~40 when arms are 130 mm long, arranged in three columns of 2-3 cirri in each radius; longest ... [details]

 Diagnosis

Cyclometra with cirri ~80 when arms are just over 100 mm long, arranged in 3 columns of 4-5 each per radial area; ... [details]

 Diagnosis

Dorometra with P2 (12 pinnulars) intermediate in size and number of pinnulars between P1 (8 pinnulars) and P3 ... [details]

 Diagnosis

Dorometra with third to fifth cirrals longest, with L/W ratio ~3. P1 10 mm long with 13 segments; P2 3.5 mm long, ... [details]

 Diagnosis

Dorometra with P3 much less than twice as long as P1 and P2, which is slightly shorter than P1. Cirri 7-11 mm long, ... [details]

 Diagnosis

Dorometra with P2 intermediate in length between P1 and P3; first three pinnules with 8-9 segments. Cirri ... [details]

 Diagnosis

Dorometra with P2 slightly smaller than P1, which is about half as long as P3, which has 18-20 segments. Cirrals ... [details]

 Diagnosis

Dorometra with P1 and P2 similar, with usually 8-10 segments, only half as long as P3, which has 13-16 segments. ... [details]

 Diagnosis

Large, robust Dorometra with P2 (13 to at least 14 pinnulars) intermediate in size and in number of pinnulars ... [details]

 Diagnosis

Eometra with centrodorsal as tall as wide at the base, which is 2.5 mm. in the type specimen. Cirri ~20, of 26-31 ... [details]

 Diagnosis

Eometra with centrodorsal basal diameter 1.33 times height. Cirri 22-27, ~18 mm long, probably of ~18 cirrals; ... [details]

 Diagnosis

A species of Erythrometra with cirri of 36-39 cirrals with arm length 60 mm. [details]

 Diagnosis

A species of Erythrometra with cirri of up to 30 cirrals and arms up to 35 mm long. [details]

 Diagnosis

Euantedon with 40-50 cirri, 10-12 mm long, with up to 15 cirrals; longest cirrals with L/W ratio up to 6 (median ... [details]

 Diagnosis

Euantedon with 15-17 cirrals; longest cirrals with L/W ratio ~4 (median width); distal cirrals decreasing in length ... [details]

 Diagnosis

Euantedon with 25 cirri of 17-26 (usually about 20) cirrals; longest cirrals with L/W ratio ~2. Arms ~40 mm long in ... [details]

 Diagnosis

Euantedon with 30 cirri, 8-10 mm long, of 13-14 cirrals; longest cirrals with L/W ratio of ~2. P1 11-15 mm long, ... [details]

 Diagnosis

Euantedon with 15-17 cirrals; longest cirrals with L/W ratio 2.0-2.5; distal cirrals only very slightly shorter; ... [details]

 Diagnosis

Euantedon with ~40 cirri with up to 25 cirrals; longest with L/W ratio 2.5-3. Arms ~100 mm long. [details]

 Diagnosis

Eumorphometra with ~40 cirri of up to 28 cirrals when arms are ~30 mm long. Brachitaxes with distinctly spinous ... [details]

 Diagnosis

Eumorphometra with ~30 cirri of up to 25 cirrals, 7 mm long when arms are ~30 mm long. Brachitaxis margins spinous; ... [details]

 Diagnosis

Eumorphometra having cirri ~45, of 34-40 cirrals, up to 12 mm long in 15 columns, when arms are 35-40 mm long. [details]

 Diagnosis

Eumorphometra with ~40 cirri (in distinct columns interradially), of 25-30 cirrals, when arms are ~35 mm long. ... [details]

 Diagnosis

Eumorphometra with ~40 cirri irregularly arranged, of 23-28 cirrals, when arm length is 25 mm. First primibrachial ... [details]

 Diagnosis

Fariometra with lateral margins of first primibrachial converging; lateral angles of axils extending beyond angles ... [details]

 Diagnosis

Fariometra with centrodorsal conical, 2.6 mm across base and 2.2 mm tall (holotype). Cirri 45-100, with up to 21 ... [details]

 Diagnosis

Fariometra with primibrachial ossicles and first two brachials in close lateral contact throughout and sharply and ... [details]

 Diagnosis

Fariometra with arms 30 mm long. Cirri ~40, of 20-22 cirrals; longest cirrals with L/W ratio ~3 (median width) with ... [details]

 Diagnosis

Fariometra with cirri of 19-21 cirrals; longest with L/W ratio 2-3 (median width). Primibrachial ossicles and first ... [details]

 Diagnosis

Large Fariometra with centrodorsal 3.5 mm across base and 3 mm tall. First primibrachials with converging lateral ... [details]

 Diagnosis

Fariometra similar to F. alcyon but with centrodorsal an equilateral triangle in lateral view, and longest cirrals ... [details]

 Diagnosis

Fariometra with second brachials shield-shaped and much longer than broad. Pinnules and cirri unknown. [details]

 Diagnosis

Fariometra with centrodorsal large, conical, twice as broad at the base as tall. Cirri ~55, with ~26 cirrals; ... [details]

 Diagnosis

Fariometra with centrodorsal conical, almost as high as wide at the base. Cirri ~60, of 21-36 cirrals; longest ... [details]

 Diagnosis

Florometra similar to F. serratissima but with third syzygy at brachials 14+15 (occasionally 12+13 to 16+17). [details]

 Diagnosis

Florometra with basal segments of oral pinnules markedly broadened but not carinate. P1 usually of ~60 segments ... [details]

 Diagnosis

Florometra with centrodorsal conical; bare aboral pole very small. Cirri ~100, of up to 38 cirrals; no aboral ... [details]

 Diagnosis

Florometra with basal pinnulars enlarged and carinate. P1 and P2 similar in length and number of segments, their ... [details]

 Diagnosis

Florometra with brachials following the third syzygy about as long as broad instead of markedly broader than long. ... [details]

 Diagnosis

Florometra with 40-77 cirri, of up to 34 cirrals; longest cirrals with L/W ratio up to 3; distal cirrals usually ... [details]

 Diagnosis

Florometra with third syzygy usually between brachials 16+17 (sometimes 15+16). Basal pinnulars of oral pinnules ... [details]

 Diagnosis

Florometra with oral pinnules slender, lacking carinate processes on basal pinnulars of proximal pinnules. Middle ... [details]

 Diagnosis

Same as genus. [details]

 Diagnosis

Same as genus. [details]

 Diagnosis

Iridometra with first three exterior (and usually first two interior) pinnules similar, elongate, longer than their ... [details]

 Diagnosis

Very large and robust Iridometra with centrodorsal large, hemispherical, with deeply concave aboral apex. Arms up ... [details]

 Diagnosis

Same as genus. [details]

 Diagnosis

Leptometra differing from L. phlangium in having cirri proportionately shorter and not tapered distally, so that ... [details]

 Diagnosis

Leptometra with centrodorsal sharply to bluntly, rounded, or truncated conical, or even somewhat columnar, usually ... [details]

 Diagnosis

Mastigometra with 50-80 cirri 12-14 mm long, of ~15 cirrals; distal cirrals with L/W ratio 1.3-1.5. P1 25 mm long ... [details]

 Diagnosis

Mastigometra with 50-90 cirri ~10 mm long, of ~16 cirrals; sixth and following cirrals with L/W ratio ~1. P1 15 mm ... [details]

 Diagnosis

Mastigometra with ~35 cirri 12-14 mm long, of 12-16 cirrals; distal cirrals with L/W ratio 1.3-1.5. P1 15 mm long ... [details]

 Diagnosis

Same as genus. [details]

 Diagnosis

Nanometra with centrodorsal conical with small aboral pole; cirri XXX-L, 30-45 (usually 35-40) segments, when arms ... [details]

 Diagnosis

Nanometra with centrodorsal broadly truncated with wide, more or less flat, aboral pole; cirri ~LXX when arms are ... [details]

 Diagnosis

Same as genus. [details]

 Diagnosis

Perometra closely resembling P. diomedeae but with all pinnules present, 13-15+ arms 50-60 mm long, less strongly ... [details]

 Diagnosis

Perometra with 10 arms only, to 90 mm long; Pa usually absent; synarthrial tubercles extravagantly developed; 35-55 ... [details]

 Diagnosis

Perometra with 10 arms (in the unique holotype) ~30 mm long; cirri XVII, up to 6 mm long with 28 segments (about a ... [details]

 Diagnosis

Perometra with cirri to 20 mm long when arm length is 60 mm, rather stout basally, with 39-47 segments; longer ... [details]

 Diagnosis

Phrixometra with cirri of up to 20 cirrals, wider distally; longest cirrals in peripheral cirri with L/W ratio ~3. ... [details]

 Diagnosis

Similar to the types of Phrixometra longipinna longipinna but with more cirri (45), fewer cirrals (17-19), more ... [details]

 Diagnosis

P. longipinna with cirri ~30, of 20-25 cirrals. Proximal pinnules all long;, P1 reaching to about brachial 12, with ... [details]

 Diagnosis

Phrixometra with cirri ~45, with <20 cirrals. P1 of ~14 segments; P2 usually first genital pinnule, similar in length and number ... [details]

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