Copepoda taxon details
Scottolana Por, 1967
|Classification: Biota > Animalia > Arthropoda > Crustacea > Multicrustacea > Hexanauplia > Copepoda > Neocopepoda > Podoplea > Harpacticoida > Canuellidae > Scottolana|
|Parent||Canuellidae Lang, 1944|
additional source Huys, R. (2009). Unresolved cases of type fixation, synonymy and homonymy in harpacticoid copepod nomenclature (Crustacea: Copepoda). Zootaxa. 2183:1-99., available online at http://www.mapress.com/zootaxa/2009/2/zt02183p099.pdf [details]|
From editor or global species databaseTaxonomy Scottolana Por, 1967 (Family Canuellidae)
Por (1967: 105) proposed the genus Scottolana to accommodate three species that had originally been placed in the genus Sunaristes Hesse, 1867: Sunaristes inopinata Thompson & Scott, 1903; S. longipes Thompson & Scott, 1903; and S. bulbosus Por, 1964a. He also considered Canuella scotti Sewell, 1940 and Sunaristes curticaudata Thompson & Scott, 1903 as potential candidate members of the genus. Subsequently, Coull (1972: 209) added Canuella canadensis Willey, 1923 and Wells (1976: 18) assigned Canuella bulbifera Chislenko, 1971 to Scottolana without making a firm recommendation for this placement. In a later review of the Canuellidae, Por (1984: 14) restricted the genus – which he puzzlingly cited as a ‘new combination’ – to just two species, Scottolana longipes (Thompson & Scott, 1903) and S. uxoris Por, 1983a, and maintained Canuella scotti as a candidate member for inclusion. He provisionally reassigned Scottolana bulbosa (Por, 1964a) to Sunaristes, proposed a new genus Coullana Por, 1984 for S. canadensis (Willey, 1923), and regarded the status of both S. curticaudata (Thompson & Scott, 1903) and S. inopinata (Thompson & Scott, 1903) as too problematic for further consideration. The genus currently includes 14 species (Mu & Huys 2004; Wells 2007).
Por (1967) did not designate a type species for the genus Scottolana, nor has any subsequent author done so. Apart from the fact that the lack of a formal type fixation made his genus-group name unavailable, it is also clear that his diagnosis of the genus did not satisfy the provisions of ICZN Art. 13.1.1. Por’s (1967: 105) definition of the genus (“... these species in which the first leg-bearing segment can be fused to the cephalothorax can be best characterized by the reduction of the armature of P IV while the other legs have still the usual armature of Canuella”) is very uninformative, to the extent that no positive assignment of a species to this genus can be made, and no characters are given that can unequivocally differentiate the genus from other taxa in the Canuellidae. Mu and Huys (2004: 2) mentioned that an application (Case 3218) had been submitted to the International Commission on Zoological Nomenclature, proposing the conservation of the generic name Scottolana Por, 1967 and the fixation of Sunaristes bulbosus Por, 1964a as type species. However, since the latter was selected on the erroneous ground that it had to be one of the originally included nominal species to be eligible for type fixation (ICZN Art. 69.2 is obviously not applicable to unavailable names; see also Art. 67.2.1), the application was subsequently withdrawn.
Given the artificial species composition of the genus Scottolana, an objective standard of reference for the application of the name is highly desirable, yet difficult to define (Mu & Huys 2004). In the interest of nomenclatural stability the generic name Scottolana (gender: feminine) is re-established here as intentionally new (ICZN Art. 16.1), taking the authorship and date of the present paper. Scottolana geei Mu & Huys, 2004 [= Scottolana geei (Mu & Huys, 2004) comb. nov.] is formally fixed as the type species (ICZN Art. 13.3), being representative of a core group of species referred to as the longipes-group (Mu & Huys 2004: 33). This group includes S. longipes (Thompson & Scott, 1903), S. longipes (Thompson & Scott, 1903) sensu Por (1964a), S. longipes (Thompson & Scott, 1903) sensu Wells (1967), S. dissimilis Fiers, 1982, S. uxoris Por, 1983a and S. longipes (Thompson Scott, 1903) sensu Wells and Rao (1987), and is characterized by at leat six apomorphies (Mu & Huys 2004: 33). In order to satisfy the provisions of ICZN Art. 13.1 a generic diagnosis is given below:
Canuellidae. Leg 1-bearing somite not fused to cephalosome. Rostrum elongate. Urosome without distinct spinule rows; hyaline frills plain. Genital double-somite of female with subcuticular ribs laterally. Female genital field large [details]
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|Citation: Boxshall, G. (2010). Scottolana Por, 1967. In: Walter, T.C. & Boxshall, G. (2017). World of Copepods database. Accessed at http://www.marinespecies.org/copepoda/aphia.php?p=taxdetails&id=534487 on 2017-03-26|