Copepoda taxon details
Pilocamptus Wells, 2007
|Classification: Biota > Animalia > Arthropoda > Crustacea > Multicrustacea > Copepoda > Neocopepoda > Podoplea > Harpacticoida > Canthocamptidae > Pilocamptus|
|Status||unaccepted (no description or definition)|
|Accepted name||Pilocamptus Huys, 2009|
|Parent||Canthocamptidae Brady, 1880|
original description Wells, J.B.J. (2007). An annotated checklist and keys to the species of Copepoda Harpacticoida (Crustacea). Zootaxa 1568:1-872., available online at http://www.mapress.com/zootaxa/2007f/z01568p872f.pdf [details]|
basis of record Walter, Chad. The world of Copepods., available online at http://www.marinespecies.org/copepoda [details]
additional source Huys, R. (2009). Unresolved cases of type fixation, synonymy and homonymy in harpacticoid copepod nomenclature (Crustacea: Copepoda). Zootaxa. 2183:1-99., available online at http://www.mapress.com/zootaxa/2009/2/zt02183p099.pdf [details]
|Species Pilocamptus africanus (Chappuis, 1933)|
Species Pilocamptus pilosus (Douwe, 1910)
Species Pilocamptus alluaudi (Chappuis, 1932) represented as Pilocamptus alluaudi (Chappuis, 1933) (alternate representation)
Species Pilocamptus kamerunensis (Kiefer, 1928) represented as Pilocamptus kamerunensis kamerunensis (Kiefer, 1928) (alternate representation)
From editor or global species databaseTaxonomy Chappuis (1929a: 43–45) fixed the types for Bryocamptus Chappuis, 1929a (i.e. Canthocamptus minutus Claus, 1863) and Echinocamptus Chappuis, 1929a (i.e. Canthocamptus echinatus Mrázek, 1893) (and thus for their respective nominotypical subgenera, ICZN Art. 67.1.1) but not for the new subgenus Arcticocamptus Chappuis, 1929a in the former or the new subgenus Limocamptus Chappuis, 1929a in the latter. Borutzky [1952: 212; 1964 (translation): 193] subsequently fixed Cyclopsine alpestris Vogt, 1845 as the type of Arcticocamptus.
Chappuis (1929a: 45) included the following species in the subgenus Echinocamptus (Echinocamptus): Canthocamptus echinatus Mrázek, 1893; C. pilosus Douwe, 1910; C. schroederi Douwe, 1915; C. georgevitchi Chappuis, 1924b; and C. kamerunensis Kiefer, 1928. In the subgenus Echinocamptus (Limocamptus) he included Canthocamptus echinatus var. luenensis Schmeil, 1894; C. hiemalis Pearse, 1905; C. hoferi Douwe, 1908; C. praegeri Scourfield, 1912; C. dacicus Chappuis, 1924a; C. calvus Brehm, 1927; and C. horai Chappuis, 1928a. Lang (1944: 28) re-assigned the subgenus Limocamptus to Bryocamptus and explicitly designated Canthocamptus echinatus Mrázek, 1893 (the type species of Echinocamptus) as the type of Bryocamptus (Limocamptus) while maintaining Echinocamptus as a valid genus (note that this taxon became hereby an “orphaned” grouping). This type fixation is invalid since C. echinatus was not originally included in the subgenus and consequently is not eligible to be fixed as the type species (ICZN Art. 67.2). The next author to fix a type species for the genus-group name Limocamptus is Borutzky [1952: 234; 1964 (translation): 214] who maintained Chappuis’s (1929a) subgeneric division of Echinocamptus and validly designated Canthocamptus echinatus var. luenensis Schmeil, 1894 as the type of Echinocamptus (Limocamptus). However, Lang (1948: 1102) claimed that Echinocamptus (L.) luenensis (Schmeil, 1894) was nothing more than a variety of Echinocamptus (E.) echinatus and that both forms certainly did not belong to separate subgenera. Except for Borutzky (1952, 1964) this synonymy was widely adopted in the freshwater copepod literature (e.g. Dussart & Defaye 1990) and effectively rendered both genus-group names Echinocamptus and Limocamptus objective synonyms of one another (ICZN Art. 61.3.2). Since they were established simultaneously, but proposed at different ranks, in the genus group, the name proposed at higher rank (Echinocamptus) takes precedence (ICZN Art. 24.1).
Since Lang (1944, 1948) had removed Canthocamptus echinatus (the type of Echinocamptus) and all species placed in Echinocamptus (Limocamptus) to a taxon of subgeneric rank in Bryocamptus, the valid name of this subgenus is Bryocamptus (Echinocamptus) and not Bryocamptus (Limocamptus) as generally accepted (e.g. Dussart & Defaye 1990; Wells 2007). The subgenus includes the following species: Bryocamptus (Echinocamptus) echinatus (Mrázek, 1893); B. (E.) hiemalis (Pearse, 1905) [and its subspecies hiemalis hiemalis (Pearse, 1905); h. verestschagini (Borutzky, 1931d); h. brevifurcatus Coker, 1934; h. yunnanensis (Borutzky, 1952); h. elongatus (Shen & Tai, 1964a); and h. yetii Löffler, 1968]; B. (E.) hoferi (Douwe, 1908); B. (E.) praegeri (Scourfield, 1912); B. (E.) dacicus (Chappuis, 1924a); B. (E.) douwei (Willey, 1925a); B. (E.) nivalis (Willey, 1925b); B. (E.) calvus (Brehm, 1927); B. (E.) horai (Chappuis, 1928a); B. (E.) morrisoni (Chappuis, 1929c) [and its subspecies morrisoni morrisoni (Chappuis, 1929c) and m. elegans (Chappuis, 1929c)]; B. (E.) smirnovi (Borutzky, 1931d); B. (E.) parvus (Borutzky, 1931d); B. (E.) viduus Kiefer, 1952; B. (E.) hostensis (Borutzky, 1972); B. (E.) pacificus Ishida, 1992; and B. (E.) lacustris Wells, 2007.
As pointed out only recently by Wells (2007: 93), Lang’s (1944, 1948) removal of the type of Echinocamptus to the genus Bryocamptus left the remaining Echinocamptus species assembled in an orphaned taxon whi [details]
[Taxonomic tree] [Google] [Google scholar] [Google images]
|Citation: Gaviria-Melo, S.; Walter, T. Chad (2013). Pilocamptus Wells, 2007. In: Walter, T.C. & Boxshall, G. (2015). World of Copepods database. Accessed at http://www.marinespecies.org/copepoda/aphia.php?p=taxdetails&id=347785 on 2016-05-31|