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Checked: verified by a taxonomic editorKliopsyllus Kunz, 1962 
AphiaID: 534149

Classification: Biota > Checked: verified by a taxonomic editorAnimalia (Kingdom) > Checked: verified by a taxonomic editorArthropoda (Phylum) > Checked: verified by a taxonomic editorCrustacea (Subphylum) > Checked: verified by a taxonomic editorMulticrustacea (Superclass) > Checked: verified by a taxonomic editorCopepoda (Subclass) > Checked: verified by a taxonomic editorNeocopepoda (Infraclass) > Checked: verified by a taxonomic editorPodoplea (Superorder) > Checked: verified by a taxonomic editorHarpacticoida (Order) > Checked: verified by a taxonomic editorParamesochridae (Family) > Checked: verified by a taxonomic editorKliopsyllus (Genus)
Status nomen nudum
Accepted name  Checked: verified by a taxonomic editorEmertonia Wilson C.B., 1932
Rank Genus
Parent Checked: verified by a taxonomic editorParamesochridae Lang, 1944
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Environment marine
Link Unreviewed: has not been verified by a taxonomic editorTo ITIS
From editor or global species database
Checked: verified by a taxonomic editorTaxonomy Kliopsyllus Kunz, 1962 (Family Paramesochridae)
As part of his revision of the family Paramesochridae Kunz (1962: 251) proposed the generic name Kliopsyllus for four species that had previously been included in other paramesochrid genera: Leptopsyllus coelebs Monard, 1928; Paramesochra holsatica Klie, 1929 (including the subspecies P. holsatica varians Kunz, 1951); Leptopsyllus constrictus Nicholls, 1935 (including the subspecies Paramesochra constricta orotavae Noodt, 1958); and Paramesochra major Nicholls, 1939b. Unfortunately Kunz (1962) failed to fix the type, rendering the genus-group name Kliopsyllus unavailable. The genus has witnessed the addition of 31 species and 5 subspecies (Wells 2007; Kornev & Chertoprud 2008), was extensively reviewed by Kunz (1981), and was regarded as a senior subjective synonym of Krishnapsyllus Kunz, 1974 by Wells et al. (1975) (even though – as a nomen nudum – it could not have entered the synonymy of other generic names). The first authorities to make the generic name Kliopsyllus available are Apostolov and Marinov (1988: 248–249) who designated Paramesochra holsatica Klie, 1929 as the type species by original designation and provided an accompanying differential diagnosis (in Bulgarian). Since the provisions of ICZN Arts 13.1.1 and 13.3 are met, the generic name should be cited as Kliopsyllus Apostolov & Marinov, 1988. However, as an available name the latter now formally becomes the junior subjective synonym of Krishnapsyllus Kunz, 1974 (type species by original designation: Krishnapsyllus furcovaricatus Kunz, 1974). None of these names can be adopted as the valid name for the genus they denote since both are also junior subjective synonyms of Emertonia Wilson, 1932 (type species by original designation: Emertonia gracilis Wilson, 1932).
Nicholls (1935: 385) listed Emertonia as a synonym of Leptopsyllus T. Scott, 1894b, while Kunz’s (1938: 228) acceptance of the former genus was conditional, depending upon the accuracy of Wilson’s (1932: 256) description of E. gracilis. In a personal communication to A. G. Nicholls (August 1937) C. B. Wilson wrote “Let me suggest that there are really three valid genera as follows, Leptopsyllus with the single species typicus, Paramesochra with the species dubia, intermedia, herdmani, robertsoni, (and) Emertonia with the species gracilis, minor, constrictus [sic], holsatica, acutata” (Nicholls 1945c: 100). Based on the different endopodal segmentation of the swimming legs, Nicholls (1945c: 103) accepted the genus as valid and included two more species, Leptopsyllus coelebs Monard, 1935a and Paramesochra pygmaea Nicholls, 1939b. His statement that “... E. pygmaea may be regarded as the genotype” has no nomenclatural validity even if Wilson (1932) had failed to fix the type species (ICZN Art. 67.5). Re-examination of Wilson’s type material in the National Museum of Natural History, Washington, D.C., led Lang (1948: 879) to rank Emertonia as a genus incertae sedis in the family Paramesochridae. Conversely, Krishnaswamy (1957: 115) followed Nicholls’s (1945c) judgement and added three new species to the genus: E. capensis Krishnaswamy, 1957, E. minuta Krishnaswamy, 1957 and E. pseudogracilis Krishnaswamy, 1957. Both Lang (1965: 377) and Kunz (1981: 11–13) rejected this generic assignment and removed all three species to Kliopsyllus. Examination by the present author of the remaining syntypes of E. gracilis deposited in the Natural History Museum, London confirmed that this species conforms entirely with Kunz’s (1962: 251; 1981: 11) and Apostolov and Marinov’s (1988: 249) diagnoses of Kliopsyllus. Consequently, the senior synonym Emertonia is reinstated here as the valid name for the genus it denotes. Its constituent species and subspecies are as follows: Paramesochra holsatica Klie, 1929 = Emertonia holsatica (Klie, 1929) comb. nov.; Emertonia gracilis Wilson, 1932 [type]; Leptopsyllus coelebs Monard, 1935a = E. coelebs (Monard, 1935a) comb. nov.; Leptopsy [details]
LSID urn:lsid:marinespecies.org:taxname:534149
Edit history
Date   action   by
2010-10-17 15:51:20Z  created  Boxshall, Geoff
2015-07-09 10:01:39Z  changed  Walter, T. Chad
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  Citation: Walter, T. Chad; Boxshall, G. (2015). Kliopsyllus Kunz, 1962. In: Walter, T.C. & Boxshall, G. (2015). World of Copepods database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=534149 on 2015-10-09
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