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Knizhin, I. B.; Antonov, A. L.; Safronov, S. N.; Weiss, S. J. (2007). New species of grayling Thymallus tugarinae sp. nova (Thymallidae) from the Amur River Basin. Journal of Ichthyology. 47(2): 123-139.
Knizhin, I. B.; Antonov, A. L.; Safronov, S. N.; Weiss, S. J. (2007). New species of grayling Thymallus tugarinae sp. nova (Thymallidae) from the Amur River Basin. Journal of Ichthyology. 47(2): 123-139.
