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Litaker, R.W., Vandersea, M.W., Faust, M.A., Kibler, S.R., Chinain, M., Holmes, M.J., Holland, W.C. & Tester, P.A. 2009. Taxonomy of Gambierdiscus including four new species, Gambierdiscus caribaeus, Gambierdiscus carolianus, Gambierdiscus carpenteri and Gambierdiscus ruetzleri (Gonyaulacales, Dinophyceae). Phycologia, 48: 344-90