Late Pliocene [details]
Sharm El Arab Member, Shagra Formation [details]
Wadi Gassus, 10 km SW Safaga, Red Sea coast, Egypt [details]
Wadi Wizr, 41km SW Quseir, Red Sea coast, Egypt [details]
Wadi Wizr, 41km SW Quseir, and Wadi Gassus, 10 km SW Safaga, Red Sea coast, Egypt [details]
HT: AUSGM E (the specimen figured on pl. 2, figs. 8-10, no number mentioned); PT: AUSGM E (the specimen figured on ... [details]
HT: AUSGM E (the specimen figured on pl. 2, figs. 1-3, no number mentioned) [details]
2 syntypes, repository unknown [details]